Cyrtodactylus calamei, Luu, Vinh Quang, Bonkowski, Michael, Nguyen, Truong Quang, Le, Minh Duc, Schneider, Nicole, Ngo, Hanh Thi & Ziegler, Thomas, 2016

Luu, Vinh Quang, Bonkowski, Michael, Nguyen, Truong Quang, Le, Minh Duc, Schneider, Nicole, Ngo, Hanh Thi & Ziegler, Thomas, 2016, Evolution in karst massifs: Cryptic diversity among bent-toed geckos along the Truong Son Range with descriptions of three new species and one new country record from Laos, Zootaxa 4107 (2), pp. 101-140 : 132-135

Cyrtodactylus irregularis group

Species. C. buchardi , C. cryptus , C. pseudoquadrivirgatus .

Characters. Adult SVL 65.0– 83.8 mm, supranasal single; dorsal tubercle rows 15–25; webbing between fingers and toes absent; tubercles on limbs present; number of precoacal and femoral pores in males 8–11; postcloacal tubercles 2–4; tail dorsally with tubercles; without enlarged subcaudals; dorsal body blotched.

Taxon

1

2 3 4 5 6 7 8 9 10 11 12 14 15 Cyrtodactylus phongnhakebangensis species group C. bansocensis 74 103.5 8‾10 8 3 0 3 0 14‾15 158‾170 86‾87 34‾35 0 16‾19 18‾21 Cyrtodactylus calamei sp. nov. * 89.3 107.5 9‾11 8‾11 3 1 4 0 10‾16 183‾193 101‾114 39‾42 0 16‾18 18‾21 C. darevskii 100 113 10‾12 9‾11 3 0 4‾5 1 16‾20 190‾216 ‾ 38‾46 0 17‾20 18‾22 Cyrtodactylus hinnamnoensis 100.6 sp. nov. * C. jaegeri 68.5 108.3 83.4 9‾12 10‾11 7‾11 3 9‾11 3 0‾1 0‾1 4‾6 5 0 1 14‾19 15‾16 179‾201 156‾164 93‾112 ‾ 35‾48 31‾32 0 1 16‾21 17‾19 18‾22 20‾23 C. jarujini 90 116 12‾16 10‾12 4 1 0 0 18‾20 169 ‾ 32‾38 0 15‾17 18‾19 C. khammouanensis 73 95 11‾12 9‾10 3 0 4‾5 0 16‾21 155‾172 ‾ 32‾38 0 18‾20 20‾23 C. lomyenensis 71.2 86.1 13‾14 11 3 0‾1 4 0 20‾24 ‾ ‾ 35‾36 0 16‾19 19‾23 C. multiporus 98 105 9‾11 9‾11 4 0 6‾8 0 16‾20 164‾181 ‾ 30‾38 0 18‾20 18‾22 C. pageli 81.8 113.2 9‾11 9 4 0 5 0 10‾14 216‾239 115‾122 41‾44 0 19‾23 19‾23 C. phongnhakebangensis 96.3 110 9‾13 8‾12 3 0‾1 3‾5 0 11‾20 161‾177 90‾107 32‾42 1 15‾20 18‾26 C. roesleri 73.5 101 10‾12 7‾10 3 0‾1 4‾5 1 13‾19 158‾187 ‾ 34‾40 0 17‾19 17‾20 C. rufford 72.5 96.8 11‾12 9‾11 3 1 4 0 14‾16 153‾167 74‾79 27‾29 0 19‾20 18‾19 Cyrtodactylus sommerladi sp. 80.3 nov.* C. soudthichaki 70.0 89.9 95.2 9‾11 10‾11 8‾9 2‾3 8‾9 3 0‾1 0‾1 5‾6 5 0 0 0‾5 19‾20 168‾192 165‾170 76‾93 78‾85 31‾39 32‾33 0 1 16‾20 16‾18 17‾24 18 C. teyniei 89.9 110 10 9 4 0 0 0 19 184 108 38 0 17‾18 19‾20 Cyrtodactylus irregularis species group C. buchardi 65 54 13‾14 10‾11 4 1 0 0 25 ‾ ‾ 30 0 14 12 C. cryptus * 83.6 73 7‾8 6‾7 4 1 0 0 15‾17 211‾218 122‾131 40‾43 0 17‾19 17‾19 C. pseudoquadrivirgatus 83.8 72.6 8‾10 7‾10 4 1 0 0 17‾18 ‾ ‾ 39‾40 1 16‾19 17‾20 Cyrtodactylus wayakonei species group C. spelaeus 91 83 9‾12 8‾10 3 1 0 0 10 156‾183 ‾ 36‾39 0 19‾20 22‾24 C. vilaphongi 86.1 68.1 9‾10 7‾9 3 0 5 0 15‾16 161‾165 106‾122 34‾36 1 18‾19 18‾20 C. wayakonei 86.8 89 7‾8 9‾10 4 0 0 0 17‾19 151‾163 85‾98 31‾35 1 17‾18 19‾20 Cyrtodactylus interdigitalis species group C. interdigitalis 80 90 10‾12 8‾9 3 1 6 0 18 174 105‾109 37‾42 1 17‾22 16‾20

13

Taxon

16 17

18 19 20 21 22 23 24 25 26 27 28 29 Reference Cyrtodactylus phongnhakebangensis species C. bansocensis 0 0 group 1 34 5–7 1 2 Greyish brοwn 1 0 1 0 0 Khammοuane Prοvince Luu et al. 2016b Cyrtodactylus calamei sp. nov. * 0 1 C. darevskii 0 0 1 1 35–39 38–44 4 4–5 0 0 2 1 Grey Brοwnish grey 1 1 1 1 1 1 0 0 0 0 Khammοuane Prοvince Khammοuane Prοvince Τhis paper Nazarοv et. al. 2014 Cyrtodactylus hinnamnoensis 0 1 sp.nov.* C. jaegeri 0 1 1 1 36–44 44 4–6 3–6 0 1 2 2 Υellοwish brοwn Greyish brοwn 1 0 0 0 1 1 0 0 1 0 Khammοuane Prοvince Khammοuane Prοvince Τhis paper Luu et al. 2014a C. jarujini 0 1 C. khammouanensis 0 0 C. lomyenensis 0 0 1 1 1 52–54 40–44 39–40 4 5–6 5 1 0 0 2 1 1 Grey brοwn Brοwn Brοwnish grey 1 0 1 1 0 0 0 1 1 0 0 0 1 1 0 Bοlikhamxay Prοvince Khammοuane Prοvince Khammοuane Prοvince Bain & Hurley 2011 Nazarοv et al. 2014 Ngο & Pauwels 2010 C. multiporus 0 1 C. pageli 0 0 C. phongnhakebangensis 0 0 1 0 1 58–60 4 32–42 5–6 4 4–5 0 1 0 2 1 2 Brοwn Light brοwn Dark grey 1 1 1 1 0 1 1 0 1 0 0 0 0 1 1 Khammοuane Prοvince vientiane Prοvince Invalid Nazarοv et al. 2014 Schneider et al. 2011 Luu et al. 2013 C. roesleri 0 0 C. rufford 0 0 1 1 20–28 42–43 5–8 4–5 1 0 2 1 Brοwnish grey Brοwn-Grey 0 1 0 0 1 1 0 0 1 0 Invalid Khammοuane Prοvince Τeynie & David 2010 Luu et al. 2016a Cyrtodactylus sommerladi sp. 0 0 nov.* C. soudthichaki 0 1 C. teyniei 0 0 1 1 1 20–26 29 – 4–6 4–5 3 0 1 1 2 2 1 Υellοwish grey Greyish brοwn Greyish brοwn 0 1 1 0 0 1 1 1 0 0 0 0 0 1 1 Khammοuane Prοvince Khammοuane Prοvince Bοrikhamxay Prοvince Τhis paper Luu et al. 2015a David et al. 2011 Cyrtodactylus irregularis species group C. buchardi 0 1 1 9 – 1 0 Dark tan 0 1 0 1 1 Champasak Prοvince David et al. 2004 C. cryptus * 0 0 C. pseudoquadrivirgatus 0 1 1 1 8–11 – 3–4 2–3 1 1 0 0 Brοwnish grey Blackish brοwn 1 1 1 1 0 0 0 0 1 1 Khammοuane Prοvince Salavan Prοvince Τhis paper Luu et al. 2013 Cyrtodactylus wayakonei species group C. spelaeus 0 1 1 8 2–3 0 1 Grey brοwn 1 1 0 0 0 vientiane Prοvince Nazarοv et al. 2014 C. vilaphongi 0 1 C. wayakonei 0 1 1 1 – 6–8 2 4 1 1 1 1 Blackish brοwn Grey-brοwn 1 1 1 1 1 1 0 0 0 0 Luang Prabang Prοvince Luang Nam Τha Prοvince Schneider et al. 2014 Nguyen et al. 2010 Cyrtodactylus interdigitalis species group C. interdigitalis 1 1 1 16–18 1–2 1 0 Dark brοwn 0 0 1 0 1 vientiane and Khammοuane prοvinces Τeynie & David 2010; οur specimens frοm Khammοuane Prοvince

Distribution. The species of this group are more widespread distributed in northern and central Laos including Khammouane, Champasak, and Salavan provinces. These species have a tree-dwelling life style.