Bicellaria farkaci, Barták, Miroslav, Plant, Adrian & Kubík, Štěpán, 2013

Barták, Miroslav, Plant, Adrian & Kubík, Štěpán, 2013, Species of Bicellaria (Diptera: Hybotidae) from Asia, Zootaxa 3710 (3), pp. 233-256 : 237-238

Bicellaria farkaci sp. nov.

( Figs 5, 6 )

Type material. HOLOTYPE ♂, China, N. Yunnan, Xue Shan nr. Zhongdian, 4200 m, 24.vi.1996, 27.49 N, 99.34 E, J. Farkač, P. Kabátek and A. Smetana (CULSP). PARATYPES: 3♂, 4♀, same data as holotype (CULSP).

Diagnosis. All setae black; third antennal segment without elongate dorsal setae; 4–6 setae on each palpus; front tibia with posteroventral setae slightly shorter than depth of tibia, but longer than ciliation; hind basitarsus narrow and hind tibia slightly swollen in both sexes. Male hypandrial processes absent, phallus with two nearly identical hooks; visible part (approximately posterior half) of female 8th sternite microtrichose.

Etymology. The species is named in honour of one of the collectors and donator of the specimens of the type series, Prof. Jan Farkač (Prague), a specialist on ground beetles ( Coleoptera , Carabidae ).

Description. Male. Head blackish-brown, brownish-grey microtrichose, black setose (including lower occiput). Holoptic, facets in dorsal half much larger than in ventral half, about 10–13 facets in line of contiguity of eyes. One pair of rather short ocellar setae. Occiput rather densely and long setose. Face slightly broadened ventrally, ventral part about 0.07 mm broad. Clypeus slightly paler grey microtrichose. Palpus short, brown, with 4–6 setae. Labrum short, brown, polished. Antenna black, 2nd segment long setose ventrally (longest setae almost 0.20 mm); 3rd segment without elongate dorsal setae; ratio of broad part of segment 3: narrow part of segment 3: arista (at 0.01 mm resolution) = 15–20: 3–5: 20–23. Thorax blackish-brown, microtrichose, mesoscutum dark brownish black in dorsal view, prescutellar area differently microtrichose than other parts of mesoscutum, appearing pale brownish grey. All thoracic setae black including fringes on squama. Chaetotaxy: 7–8 narrowly biserial acrostichals; 8–9 uniserial dorsocentrals; 1 intrahumeral, 1 posthumeral and 0–2 additional setae in posthumeral and intrahumeral areas; 1 long and 1–2 shorter postpronotals; notopleuron with 3–4 unequally long setae arranged in one irregular row; 1 supraalar and 1 strong prealar; 1 long postalar; 3 pairs of long scutellars; pleura, antepronotum and sternites without any setae. Legs including coxae black, microtrichose and black setose. Fore femur with row of very short anteroventrals and row of longer posteroventrals (slightly shorter than depth of femur). Fore tibia with 0–2 dorsal and 0–1 submedian posterodorsal setae about 1.5X as long as depth of tibia, otherwise short setose, posteroventrals in apical third about as long as depth of tibia, pilosity apparently absent (similar to B. spuria ); conspicuous anteroventral sense organ ovoid, about 0.10 mm long, consisting of central area of very short hairs and surrounding ovoid circle devoid of microtrichiae. Mid femur with sparse row of anteroventrals one-third as long as depth of femur and with much stronger and longer posteroventrals slightly longer than depth of femur. Mid tibia with 2 anterodorsal (1 subbasal and 1 submedian) and 1 submedian posterodorsal setae 2X as long as depth of tibia and 2 ventral preapical setae. Hind femur with 15–17 anterodorsal setae on basal third slightly longer than depth of femur, with row of 18–20 anteroventrals about as long as depth of femur, much shorter posteroventrals absent on apical half to third. Hind tibia very slightly swollen in apical part (0.12 mm at broadest point in comparison with 0.08 mm in basal part) being much narrower than hind femur (0.18 mm at broadest point), both dorsal and ventral setae slightly longer than depth of tibia, several anterodorsals up to 0.20 mm long. Tarsi of all legs slender, short setose. Wing slightly brownish infuscated; stigma dark brown, long (0.80 mm), almost parallel-sided and narrow (0.05 mm), much narrower than cell r1 (about 0.13 mm broad at narrowest point). Distance between tips of veins R1 and R2+3 about 0.40 mm. Squama dark brown, halter dark brown. Abdomen brown, microtrichose, entirely black setose, dorsally with short setae, marginals on sides of tergites 2–5 slightly longer than corresponding segments, on subsequent tergites slightly shorter. Genitalia ( Figs 5, 6 ) with hypandrial processes absent, distal part of hypandrium with some 5 long setae on each half; postgonites triangle-shaped, two nearly identical phallic hooks present. Female. Tergites entirely pruinose, sternite 8 microtrichose on apical (visible) half, polished on basal half. Length: body 2.7–3.5 mm, wing 3.2–4.0 mm.

Remarks. All specimens of the type series are rather damaged, devoid of most mesoscutal setae; however, their arrangement, number and length may be judged from their articulations and several setae present on some specimens. The new species is quite different from all known species of Bicellaria in having hypandrium rounded caudally, without processes. Also the combination of setose palpus with third antennal segment lacking long dorsal setae is not present in any known species and the microtrichose female 8th abdominal sternite is also a very peculiar feature. According to all these characters this species is easily recognizable from all other species of Bicellaria worldwide.

Figures

FIGURES 1 – 8. Hypandrium (ventral view) and phallus with postgonites (lateral view). 1 – 2. Bicellaria amankutanensis sp. nov. 3 – 4. B. chimganensis sp. nov. 5 – 6. B. farkaci sp. nov. 7 – 8. B. femorata Collin. Scale bars = 0.10 mm.