Geostiba (Tropogastrosipalia) extensicollis, Assing, 2010

Assing, V., 2010, A revision of Geostiba of the Western Palaearctic region. XX. Four new species from Turkey and Albania, and additional, Linzer biologische Beiträge 42 (2), pp. 1125-1138 : 1132-1135

Geostiba (Tropogastrosipalia) extensicollis nov.sp. ( Figs 15-21 )

T y p e m a t e r i a l: Holotype: "N 39°50'08 E 28°25'31 (19), TR Bursa, 16.4.2010, Devecikonagi, 640 m, leg. Brachat & Meybohm / Holotypus Geostiba extensicollis sp. n. det. V. Assing 2010" (cAss) . Paratypes: 1, 1: same data as holotype (cAss) ; 1: "N 39°54'49 E 28°28'26 (15), TR Bursa, 15.4.2010, Karaoman [recte: Karaorman ], 440 m, leg. Brachat & Meybohm " (cAss) ; 1, 1: "TR - Bursa (15), S Mustafa Kemalpaúa, 7 km SW Karaorman , 440 m, 15.IV.2010 / N 39°54'49,3 E 28°28'26,7, leg. Meybohm & Brachat " (cAss) .

D e s c r i p t i o n: Body length 2.3-3.1 mm. Habitus as in Fig. 15 . Coloration variable: head reddish-brown to blackish-brown, usually somewhat darker than pronotum; pronotum reddish-brown to dark-brown; elytra reddish to reddish-brown; abdomen blackish-brown, usually with the anterior segments and the apex more or less extensively paler brown; legs dark-yellowish; antennae reddish-brown, with the basal 2-3 antennomeres reddish.

Head weakly oblong ( Fig. 16 ); punctation extremely fine, barely noticeable; microsculpture shallow, but distinct. Eyes relatively large, more than half the length of postocular region in dorsal view ( Fig. 17 ).

Pronotum approximately 1.2 times as wide as head ( Fig. 16 ), with pronounced sexual dimorphism; punctation extremely fine, barely noticeable; microreticulation similar to that of head or slightly more pronounced.

Elytra with pronounced sexual dimorphism, less than half as long as pronotum ( Fig. 16 ). Hind wings absent.

Abdomen wider than elytra; punctation with sexual dimorphism, moderately dense on anterior tergites and sparse on posterior tergites; interstices with distinct microsculpture; posterior margin of tergite VII with very narrow rudiment of a palisade fringe; anterior tergites with or without, tergite VII with sexual dimorphism; posterior margin of tergite VIII convex in both sexes.

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(with fully developed secondary sexual characters): pronotum oblong, approximately 1.2 times as long as broad, posteriorly produced and concealing scutellum, posterior margin truncate to indistinctly concave in the middle, lateral margins posteriorly not sinuate, posterior angles not marked ( Fig. 16 ); elytra broadly transverse, 1.25-1.30 times as wide as pronotum, with narrow and moderately elevated sutural carinae extending from apex of scutellum almost to posterior margin of elytra, punctation granulose, interstices with very weak, barely noticeable microsculpture, disc rather deeply and extensively impressed; abdomen with moderately fine punctation; anterior abdominal tergites unmodified or with indistinct median elevations on tergites IV or III-IV; tergite VII at posterior margin with rather short, stout, suberect (lateral view), wedge-shaped (anterodorsal view) spine-like process ( Fig. 18 ); posterior margin of sternite VIII convex; median lobe of aedeagus with cristal process of somewhat variable shape ( Figs 19-20 ).

: pronotum approximately as wide as long, posterior margin regularly and weakly convex; elytra less broad and less transverse than in, approximately 1.2 times as wide as head, punctation fine and non-granulose, interstices with distinct microreticulation; abdomen with very fine punctation; sternite VIII with broadly convex posterior margin; spermatheca as in Fig. 21 .

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I n t r a s p e c i f i c v a r i a t i o n: In the three males examined, the degree of intraspecific variation is relatively low. In one of the male paratypes, the pronotum is slightly narrower and less produced posteriorly and the sutural carinae are slightly shorter, but in other respects the secondary sexual characters are similar to those of the holotype.

E t y m o l o g y: The specific epithet (Latin, adjective) alludes to the conspicuously long male pronotum.

C o m p a r a t i v e n o t e s: InthekeyinASSING (2009), the species would key out either at couplet 82 together with G. heliophila ASSING from Kastamonu or at couplet 87 together with G. aydinica ASSING from the Aydın Da÷ları, depending on whether or not median elevations are present on the anterior tergites. In order to account for G. extensicollis , the key is modified as follows:

78: abdominal tergites III and/or IV each with smooth subcircular tubercle near anterior impression. ( G. biformis and G. extensicollis , in which tergites III and IV may or may not have indistinct tubercles, will key out in both alternatives.).......................................79 -: abdominal tergites III and IV unmodified....................................................................84 82: pronotum gradually tapering posteriad, i.e., lateral margins regularly converging, not sinuate near posterior angles ( Fig. 16 ). Species from northern or northwestern Anatolia...........................................................................................................................82a -: pronotum abruptly narrowed posteriad, lateral margin at least weakly sinuate near posterior angles; process of tergite VII longer, more erect, and distinctly stouter. Species from southern Anatolia........................................................................................83 82a Body smaller and of more slender habitus ( ASSING 2009: Figs 9-10 ).: pronotum relatively smaller, posteriorly usually distinctly concave in the middle, and less oblong, 1.10-1.15 times as long as broad ( ASSING 2009: Figs 9-10 ); elytra less broad, approximately 1.5 times as wide as pronotum; sutural carinae shorter and less elevated ( ASSING 2009: Figs 9-10 ); tergite IV with smooth median tubercle ( ASSING 2009: Figs 9-10 ); process of tergite VII short, acute, and suberect ( ASSING 2009: Fig. 11 ); median lobe of aedeagus with longer and more slender cristal process ( ASSING 2009: Fig. 13 ).

: spermatheca with proximal portion of capsule conspicuously transparent ( ASSING 2009: Fig. 14 ). Kastamonu .................................................................. G. heliophila ASSING - Body on average larger and broader ( Fig. 15 ).: pronotum relatively larger, posteriorly truncate or very weakly concave, and more oblong, approximately 1.2 times as long as wide; elytra broader and more transverse, 1.25-1.30 times as wide as pronotum ( Fig. 16 ); sutural carinae more pronounced and almost reaching posterior margin of elytra ( Fig. 16 ); tergite IV with or without very indistinct median elevation; process of tergite VII stouter ( Fig. 18 ); median lobe of aedeagus with shorter cristal process ( Figs 19-20 ).: proximal portion of spermathecal capsule not conspicuously transparent. Northwestern Anatolia: Bursa ..................................... G. extensicollis nov.sp.

87: pronotum (in large) more oblong (up to 1.25 times as long as wide) and more strongly projecting posteriorly ( Fig. 16 ; ASSING 2006: Fig. 46); elytra with more strongly elevated and long sutural carina extending to posterior elytra margin (or nearly so), with pronounced impressions, and with coarser and more distinctly granulose puncturation ( Fig. 16 ; ASSING 2006: Figs 46, 48). Species from western or northwestern Anatolia.....................................................................................................87a

-: pronotum much less oblong and less strongly projecting posteriorly; elytra with shorter and less elevated sutural carinae not reaching posterior elytral margin, impressions, if present, shallower.....................................................................................88

87a Forebody with less pronounced microsculpture and more glossy.: process of tergite VII more erect, anteriorly somewhat flattened, and in antero-dorsal view more slender and apically rounded ( ASSING 2006: Figs 49-50); median lobe of aedeagus smaller, approximately 0.28 mm long and shaped as in ASSING (2006: Figs 49-51).: proximal end of spermathecal capsule longer and more slender ( ASSING 2006: Fig. 52). Western Anatolia (Aydın): Aydın Da÷ları ( ASSING (2006: Map 3) .................... G. aydinica ASSING

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- Forebody with more pronounced microsculpture and less shiny ( Fig. 16 ).: process of tergite VII suberect, anteriorly convex in cross-section, and in antero-dorsal view wedge-shaped and apically acute; median lobe of aedeagus larger, approximately 0.32 mm long and shaped as in Figs 19-20 .: proximal end of spermathecal capsule bulbous and shorter ( Fig. 21 ). Northwestern Anatolia: Bursa .......... G. extensicollis nov.sp.

D i s t r i b u t i o n a n d b i o n o m i c s: The type specimens were collected in two localities in the mountain range to the south of Mustafakemalpaúa in Bursa province, northwestern Anatolia, at relatively low altitudes (440 and 640 m). They were sifted from leaf litter in beech forests (MEYBOHM pers. comm.).

Figures

Figs 15-21: Geostiba extensicollis nov.sp. (15-19: holotype): (15) male habitus; (16) male forebody; (17) head in lateral view; (18) male abdominal segments VI-VIII in lateral view; (19-20) median lobe of aedeagus in lateral view; (21) spermatheca. Scale bars: 15: 1.0 mm; 16-18: 0.5 mm; 19-21: 0.1 mm.

Figs 7-14: Geostiba kazica nov.sp. (7-8, 10-11: holotype): (7) male habitus; (8-9) male forebody; (10) head in lateral view; (11) male abdominal segments VI-X in lateral view; (12-13) median lobe of aedeagus in lateral view; (14) spermatheca. Scale bars: 7: 1.0 mm; 8-11: 0.5 mm; 12-14: 0.1 mm.