Anomala pincelada Filippini , Galante

Filippini, Valentina, Galante, Eduardo & Micó, Estefanía, 2015, Description of six new species of Anomalini from Costa Rica (Coleoptera: Scarabaeidae: Rutelinae), Zootaxa 3948 (1) : -

publication ID	https://doi.org/ 10.11646/zootaxa.3948.1.2
publication LSID	lsid:zoobank.org:pub:9A0330AD-EEF7-4A80-8D80-1B6116BD856F
DOI	https://doi.org/10.5281/zenodo.6122706
persistent identifier	https://treatment.plazi.org/id/03E1981E-B53A-FFE9-FF59-FE2C0295EF65
treatment provided by	Plazi
scientific name	Anomala pincelada Filippini , Galante
status

Treatment

Anomala pincelada Filippini, Galante , & Micó, new species

Fig. 4 View FIGURES 1 – 6

Material examined. Holotype: ♂"Fca. Jenny, 30 km N de Liberia, P.N. Guanacaste, Prov. Guan., COSTA RICA. 18-25 Abr 1993, E. Araya, L- N 316200_364400 / INBIOCRI001167801" deposited in INBIO .

Paratypes (9): 1♂ "3 Km este de Cuajiniquil, 300m, Prov. Guanacaste, Costa Rica, 25 jun 1992, III curso Parataxon. L-N 325600_355200 / INBIOCRI000863924"; 1♂ "Santa Rosa National Park Guanacaste Prov. COSTA RICA. 9-11 May 1980 DH Janzen & W Hallwachs / INBIOCRI001116980"; 1♂, 1♀ "Fca. Jenny, 30 km N de Liberia, P.N. Guanacaste, Prov. Guan., COSTA RICA. 9-14 May 1993. E. Araya, L-N 316200_364400" INBIOCRI001183402 and INBIOCRI001183321; 1♀ "Tierras Morenas, Rio San Lorenzo, Tenorio, Prov. Guana, COSTA RICA, 1050m. May 1993. G. Rodriguez, L S 283950_424500 # 2118 / INBIOCRI001180936"; 1♂ "Sector Las Pailas, P. N. Guanacaste, A. C. Guanacaste, Prov. Guana, COSTA RICA. 800 m. 6-26 Jun 1994, K. Taylor, L N 309500_389500 # 3063 / INBIOCRI001909257"; 2♀ "Finca Jenny, 30 km N de Liberia, P.N. Guanacaste, Prov. Guan., COSTA RICA. 18-25 Abr 1993, E. Araya, L- N 316200_364400" INBIOCRI001167798 and CEUA 00106161; 1♂ "Est. Murcielago, 8 km S. O. de Cuajiniquil, Prov. Guana, COSTA RICA. 100m. 16 Jun- 4 Jul 1993. F. Quesada, L N 320300_347200 # 2177 / CEUA 00106160".

Holotype male description. Body shape elongate. Length 13.2 mm. Width 7.9 mm.

Color. Head reddish brown. Pronotum ochre with an irregular median brown macula and small maculae on lateral foveae. Scutellum dark brown. Elytra ochre with a dark brown macula at each side of scutellum and dark stripes on apical third of first interstice, from basal to apical calli and near lateral margins. Pygidium ochre. Legs ochre with base and apex of tibiae dark brown; protarsi and mesotarsi reddish brown, metatarsi dark brown. Venter ochre.

Clypeus trapezoidal, with anterior angles widely curved; surface densely punctate-reticulate, flat. Anterior margin straight, clypeus anteriorly thin. Clypeus ratio width/length 2.1. Frons densely punctate, flat. Ocular canthum long, thin, with acute apex. Interocular ratio (interocular width/width of eye): 2.6. Antenna: ratio funiculus/club 0.8.

Pronotum subtrapezoidal, ratio width/length 1.7, lateral margins regularly convex. Anterior angles quadrate and blunt. Posterior angles obtuse and blunt. Basal margin sinuate, subapical bead complete. Surface with fine, dense punctures.

Scutellum. Shape subtriangular, with rounded sides and blunt apex. Ratio width/length 1.4. Surface densely punctate.

Elytra with costae defined by regular rows of punctures, subsutural interstices with 2–3 rows of punctures; maximum width towards apex.

Pygidium with coalescing punctures and long setae at apical margin. Ratio width/length 1.6.

Space between the mesocoxae narrow, flat. Mesometasternal suture well defined, at base of mesocoxae. Abdominal sternites with 1 row of setae and 2–3 rows of sparse, fine punctures per sternite. Terminal sternite punctate with subapical bead narrow and sinuate, apical membrane wide.

Protibia (Fig. 10) tridentate; apical tooth long and curved; middle tooth below internal apex of protibia, triangular in shape, acute. Metatibia stout, fusiform, ratio length/width 3.0. First external carina consists of a row of sparse setae, surface punctate above second external carina and rugose below. Protarsal claws: internal claw bifurcate, with upper branch of same length and half the width of the lower branch. Inferior margin with a sharp angle near base.

Aedeagus ( Fig. 16 View FIGURES 13 – 18 ): parameres deeply and widely sinuate anteriorly in a lateral view, becoming nearly bifurcate. Endophallus ( Fig. 22 View FIGURES 19 – 22 ) with 1 long lateral sacculus tapering at apex; ejaculatory duct opening on the other side, at base; 1 small diverticle with long spines ventrally at base.

Variation. Macula on pronotal disc of different size, from a subpentagonal macula not reaching posterior margin to covering most of the surface with ochre sides; a median narrow, faint, lighter band may be present; maculae on elytra of variable length; metatibia from ochre to red with darker apex. Female similar to male, protibia (Fig. 10) with longer and wider apical tooth; internal protarsal claw with curved inferior margin.

Body length 13.2–16.1 mm, body width 7.9–9.2 mm. Clypeus w/l: 2.0–2.3. Interocular ratio (interocular width/width of eye): 2.5–2.9. Antenna: ratio funiculus/club 0.7–0.8. Pronotum w/l: 1.7–1.8. Scutellum w/l: 1.4– 1.6. Pygidium w/l: 1.5–1.6. Metatibia l/w: 2.5–3.0.

Diagnosis. Externally nearly identical to A. inconstans Burmeister, 1844 ( Colombia, MLUH), it is separated from this species by the shape of male genitalia, which has bifurcated apex of parameres, whereas A. inconstans has parameres with thin and curved apex, blunt and wide ventral angle, and a frontal lobe-like expansion on anterior part of ventral margin ( Fig. 33 View FIGURES 27 – 33. 27 ). Specimens from Mexico, Paraguay, Guatemala, and Venezuela from MNHUB collection, also similar to A. inconstans , all show different aedeagi, so these species are part of a large and widely distributed species complex.

Etymology. Latinized from Spanish noun pincelada , brush stroke, for the elytral pattern. This name should be treated as a noun in apposition.

Distribution. Guanacaste mountain range, from 200–600 m ( Fig. 35 View FIGURE 34 – 37 ).