Galapagonotus, 2000

ANDERSON, ROBERT S. & LANTERI, ANALIA A., 2000, New Genera and Species of Weevils from the Galapagos Islands, Ecuador, and Cocos Island, Costa Rica (Coleoptera; Curculionidae; Entiminae; Entimini), American Museum Novitates 3299, pp. 1-16 : 3-13

New World Entimini rather than with Rhyncogonus .

GALAPAGONOTUS ANDERSON AND LANTERI , NEW GENUS Figures 1–9

TYPE SPECIES: Otiorhynchus cuneiformis Waterhouse, 1845: 38 , here designated.

ETYMOLOGY: This genus is named for the Galapagos Islands.

DIAGNOSIS: Body length 4.8–7.5 mm. Vestiture of flat scales and fine erect setae, setae longest on elytra. Mandibles with interior cutting edge lacking or very slightly developed basally, with numerous setae around periphery of scar and along ventral surface. Antennal scape with dense, round appressed scales; in repose, passing over middle of eye. Metepisternal suture present; metepisternum broad. Femora simple, lacking tooth. Metatibia with apical bevel broad, glabrous; apical comb of setae slightly ascended along outer margin of tibia; mucro single, apical margin of tibia not excised adjacent to base of mucro; tarsal groove squamose. Tarsal claws free, lacking basal tooth. Male with aedeagus cylindrical, apex not reflexed, apical setae lacking. Female with sternite 8 flat, not keeled, subtriangular in shape; hemisternites in dorsal view separate throughout length; styli present.

IDENTIFICATION: We know of no similar genus with which Galapagonotus could be confused. Within the weevil fauna of the Galapagos, this genus is easily identified by the short, broad rostrum lacking a median sulcus; mandible with prominent scar; and antennal scape in repose lying over the middle of the eye.

DISTRIBUTION AND DIVERSITY: This genus is endemic to the Galapagos Islands of Ecuador. Although Franz (1985) noted the presence of two species on the Galapagos Islands, one of which was found on San Cristobal Island, the other on Santa Cruz Island, we recognize only one species, Galapagonotus cuneiformis (Waterhouse) , as present on the archipelago. We have not been able to recognize patterns of variation among islands that warrant recognition of more than one species .

DESCRIPTION: Body length male 4.8–6.5 mm, female 5.5–7.5 mm. Cuticle dark reddish brown to black. Vestiture of round to tear-drop shaped scales with slight metallic reflection; scales moderately dense to dense dorsally and sparser ventrally on legs. Dorsally and ventrally also with scattered, elongate fine setae; setae longest on elytral disk.

Head with rostrum short, broad; flattened dorsally, widest at apex. Pterygia (throughout length) and scrobes (at apex) visible in dorsal view. Scrobes well defined at point of antennal insertion, vaguely defined and open posteriorly immediately in front of eye. Epistoma with raised anterior margin, moderately emarginate medially. Mandible with large prominent scar (indicating point of attachment of cusp) and numerous long, curved setae surrounding scar and along ventral margin; interior cutting edge lacking or slightly developed basally. Prementum trapezoidal, broad, widest at apex, with single pair of elongate setae at anterolateral angle; labial palpi not visible in ventral view. Eyes laterally to very slightly dorsolaterally situated, rounded to slightly elongate-oval, very convex and prominent. Head not constricted behind eyes. Antenna with scape elongate, reaching anterior margin of pronotum, in repose passing over middle of eye; with dense, rounded appressed scales and erect hairlike scales. Antennal funicle of seven articles; article 1 elongate, slightly shorter than, to subequal in length to, article 2; articles 3–7 much shorter, each about ½ length of article 2, very slightly longer than wide; articles 1– 7 with elongate appressed, hairlike scales in addition to sparse, erect hairlike scales; appressed hairlike scales densest on article 1 and 2. Antennal club elongate-oval, setose, composed of three articles.

Pronotum cylindrical, in dorsal view widest at middle, with dense round appressed scales obscuring underlying cuticle, and scattered fine erect setae. Postocular lobes absent, anterolateral margin of pronotum more or less straight.

Elytra as wide as pronotum at base, humeri rounded; striae (1–9 complete, 10 short) not deeply impressed, punctures of striae moderately large and moderately deep. Vestiture of moderately dense, appressed, teardrop shaped scales; also of scattered, elongate fine, erect setae; spacing of scales exposing underlying cuticle. Scutellum visible, triangular, glabrous. Hind wings lacking.

Legs elongate, with vestiture of rounded appressed scales and scattered fine erect setae; setae primarily arranged along inner margin of femora and tibiae. Femora clavate, widest at apical ⅓ simple, lacking tooth. All tibiae more or less straight; inner margins with small to minute rounded asperities in apical ½; mucro moderately large, curved, smaller on meso- and metatibiae. Metatibia with apical bevel broad, glabrous; apical comb of setae slightly ascended along outer margin of tibia; mucro single, apical margin of tibia not excised adjacent to base of mucro; mucro larger on male than on female. Tarsal groove squamose, with from one to a few large appressed scales. Tarsi elongate, article 1 slightly longer than 2, article 2 slightly longer than 3, article 3 bilobed, article 4 elongate, extended beyond apices of 3 by slightly greater than length of article 3; ventrally with elongate, moderately dense, fine, wispy vestiture on articles 1 and 2, vestiture slightly denser on article 3; claws simple, divergent, free.

Prosternum with procoxae contiguous, situated slightly closer to anterior margin than to posterior margin. Mesocoxae proximate, separated by more or less ¼ diameter of mesocoxa. Mesosternum sparsely punctate, with scattered sparse rounded scales. Mesepimeron short-trapezoidal, anterior margin direct- ed to elytra then turned anteriorly such that mesepisternum contacts elytron in a strip near extreme base of elytron. Metasternum short, concave medially; vestiture of moderately dense, appressed, tear-drop shaped scales and sparse fine erect setae. Metepisternal suture present, distinct and deep in anterior ½, indistinct, not impressed in posterior ½; metepisternum broad, 5–6 times as long as wide. Metacoxae widely separated by about ⅔ diameter of a metacoxa.

Abdomen with scattered appressed teardrop shaped scales and sparse, fine erect setae. Visible sternite 1 very slightly longer than 2; 3 and 4 subequal in length, short, their combined length slightly less than ⅔ length of visible sternite 2; visible sternite 5 longer than length of 3 and 4 combined. Base of visible sternite 1 concave in male, flat to slightly convex in female. Apex of visible sternite 5 very slightly emarginate at middle in male; rounded in female. Tergite 7 transverse in male, with posterior margin broadly emarginate at middle, posterolateral angles projected; elongate in female, with posterior margin narrowly truncate.

Male genitalia. Sternite 8 large, trapezoidal, posterior margin broadly emarginate, not cleft; sternite 9 long, broad; Tegmen lightly sclerotized, parameres developed, directed anteriad, very lightly sclerotized. Aedeagus cylindrical, sclerotized throughout; apex not reflexed, apical setae lacking. Apodemes subequal in length to aedeagus.

Female genitalia. Sternite 8 small, flat, subtriangular, longer than wide, widest at base; with pair of more heavily sclerotized, slightly divergent lines from base to midlength; apical ⅓ with elongate, erect setae; apodeme approximately 1.33 times length sternite. Ovipositor relatively short, less than ½ length of abdomen, lacking setae; baculi absent; hemisternites sclerotized, short, subdivided into apical and basal portions, apical portion less than ½ length of basal portion; in dorsal view separate throughout length; styli distinct, elongate. Spermatheca subcylindrical, point of insertion of duct not developed, point of insertion of gland globose, proximal.

Galapagonotus cuneiformis (Waterhouse) , new combination Figures 1–9

Otiorhynchus cuneiformis Waterhouse, 1845: 38 . Waterhouse, 1877: 82. Linell, 1898: 267.

Amphideritus cuneiformis ; Van Dyke, 1953: 142.

[no assigned genus] cuneiformis ; Kuschel, 1986: 67 ( Barynotini , incertae sedis).

TYPES: As noted by Franz (1985) the type of Otiorhynchus cuneiformis Waterhouse is missing and a neotype must be designated. Neotype male, dissected, here designated, labeled. ‘‘ECU: Galapagos /Puntudo, Scalesia / 650m, 1–8.iv.89/FIT, S.Peck, 89-199,’’ with genitalia vial and our designation label (BMNH).

DESCRIPTION: Body length male 4.8–6.5 mm, female 5.5–7.5 mm; body width male 2.3–3.0 mm, female 2.4–3.2 mm. Scales white, light to dark brown or green, usually with slight metallic reflection, not forming distinct elytral pattern. Head with rostrum irregularly, densely punctate to distinctly rugose in basal ½ to ⅔; very sparsely and finely punctate in apical ½ to ⅓. Epistoma emarginate medially in broad V shape, laterally with three long, curved setae per side. Pronotum with surface sculpture slightly irregular, distinct punctures visible, fine and deep, irregularly spaced. Elytra gradually expanded posteriorly to posterior ⅔ then attenuate to apex; intervals flat except for bases of intervals 3 and 5, which are slightly elevated; erect setae of intervals arranged irregularly in multiple rows. Aedeagus moderately curved; apex slightly produced into narrowly subacuminate tip; apical and basal regions slightly expanded in dorsal view, slightly wider than intervening length. Internal sac visible at base of aedeagus, slightly protrud- ed, with transverse apical sclerite complex.

DISTRIBUTION: ECUADOR. Galapagos Islands. Floreana Island. 300 m, 15.II.1964, G . Kuschel ( NZAC, 1) . Santiago Island. Aguacate Camp. 550 m, mossy forest, FIT , 7–13.IV.1992, S . Peck ( AMNH, 2; CMNC, 4). Aguacate (1 km NE), 600 m, 4–9. VI .1991, humid forest FIT, S. Peck ( CMNC, 1) . San Cristobal Island. [as Chatham Island]. July 1906, F . X. Williams ( CASC, 1); January 24–30, 1906, F . X. Williams ( CASC, 1). Poza Colorada , 550 m, sweeping, 19.III.1996, S . Peck ( CMNC, 1). Gebirge b. Progreso, V .–VI.1975, H. Franz ( NZAC, 2) . Santa Cruz Island. Cerro Crocker subtop, 790 m, fern sedge formol traps, 10 – 30.IV.1996, S . Peck ( CMNC, 1). Puntudo , 700 m, pampa zone shrub litter, 2.II.1989, S . Peck ( CMNC, 1). Puntudo , 650 m, Scalesia forest FIT , 1–29.II.1989, S . Peck & B . Sinclair ( CMNC, 4). Puntudo (1 km N), 650 m, Scalesia forest FIT , 1–8.IV.1989, S . Peck ( CMNC, 2). Wald über Santa Rosa , V .– VI ., 1975, H. Franz ( NZAC, 1) .

NATURAL HISTORY: As far as is known, this species is native and endemic to the Galapagos Islands. Specimens have been collect- ed on four islands and generally at upper elevations from 300 to 790 m in native Scalesia, Miconia , and fern-sedge habitats (see Peck and Kukalova-Peck, 1990: 1620 for discussion of habitats). Adults lack functional hind wings. No details are known of food habits; most broad-nosed weevils are general foliage feeders as adults, and root feeders as larvae.

COCONOTUS ANDERSON AND LANTERI , NEW GENUS Figures 10–41

TYPE SPECIES: Coconotus williamsi Anderson and Lanteri , by present designation.

ETYMOLOGY: This genus is named for Cocos Island.

DIAGNOSIS: Body length 5.5–8.1 mm. Vestiture of flat, round, often metallic scales; fine setae present only on elytral declivity. Mandibles with inner cutting edge well developed, bladelike, with prominent inwardly directed tooth, with numerous setae around periphery of scar and along ventral surface. Antennal scape with only fine appressed setae, in repose, passing over extreme lower portion of eye or under eye. Metepisternal suture present: metepisternum extremely nar- row. Humeri with short, distinct ridgelike keel in female, evenly rounded in male. Metatibia with apical bevel broad, glabrous; apical comb of setae not ascended along outer margin of tibia; mucro single, but apical margin of metatibia excised adjacent to base of mucro such that a second tooth is evident on the margin at the apex of the excision; tarsal groove squamose. Tarsal claws free, lacking basal tooth. Male with aedeagus cylindrical, apex reflexed, apical setae lacking. Female with sternite 8 flat, not keeled, subrhomboidal in shape; hemisternites in dorsal view fused at apex, styli absent.

IDENTIFICATION: Coconotus keys to couplet 34 in the Barynotini portion of the key to world Brachyderinae of Emden (1944). Coconotus is easily distinguished from other taxa that key to this point by the absence of scales on the antennal scape and by the keeled humeri in females.

DISTRIBUTION AND DIVERSITY: This genus is endemic to Cocos Island of Costa Rica. Three species, Coconotus williamsi Anderson and Lanteri , C. kuscheli Anderson and Lanteri , and C. tuberculatus Anderson and Lanteri are described herein. Hogue and Miller (1981) noted a taxon ‘‘genus near Epicaerus , ’’ which is likely Coconotus ; however, we have been unable to locate representative specimens.

DESCRIPTION: Body length male 5.5–8.1 mm, female 6.1–6.8 mm; width male 2.4– 4.0 mm, female 2.8–3.0 mm. Cuticle dark reddish brown to black. Vestiture of round scales, each with some degree of metallic reflection (scales may be greasy and appear black in some specimens), not forming distinct pattern; moderately dense to dense dorsally and on legs, sparser ventrally. Ventrally and dorsally on apical declivity of elytra also with scattered, elongate, fine setae.

Head with rostrum short, broad; flattened dorsally, widest at apex. Pterygia (throughout length) and scrobes (at apex) visible in dorsal view. Scrobes well defined at point of antennal insertion, poorly defined dorsally, but well-defined ventrally; somewhat open dorsally, but with deeper channel directed below eye. Epistoma with flat to very slightly raised anterior margin, very slightly emarginate medially, laterally with two long, curved setae per side. Mandible with small scar (indicating point of attachment of cusp), with numerous setae around periphery of scar and along ventral surface; interior cutting edge well developed, bladelike, with prominent inwardly directed tooth. Prementum trapezoidal, broad, widest at apex, with single pair of elongate setae at anterolateral angle; labial palpi visible in ventral view. Antenna with scape having dense, fine setae only; elongate, reaching anterior margin of pronotum; in repose passing over extreme lower portion of eye or under eye. Antennal funicle of seven articles: article 1 elongate, subequal in length to article 2; articles 3–7 much shorter, each about ½ length of article 2, very slightly longer than wide; articles 1– 7 with elongate, appressed hairlike scales in addition to sparse erect vestiture. Antennal club elongate-oval, setose, composed of three articles. Frons slightly concave. Eyes large, laterally to very slightly dorsolaterally situated, rounded, convex, and prominent. Head not constricted behind eyes.

Pronotum cylindrical, in dorsal view widest at middle, with dense, round appressed scales. Postocular lobes lacking, anterolateral margin of pronotum more or less straight.

Elytra as wide as pronotum at base, gradually expanded posteriorly to midlength then attenuate to apex, broadly flattened in male, less so in female; striae (1–9 complete, 10 short) not, to moderately impressed, punctures of striae moderately large and moderately deep, serially arranged or scattered. Humeri rounded, with short, distinct ridgelike keel in female, lacking in male. Vestiture of moderately dense, appressed, round scales; scales in most places obscuring underlying cuticle; fine, erect setae only visible on elytral declivity. Scutellum visible; small, triangular, glabrous. Hind wings lacking.

Legs elongate, with vestiture of rounded appressed scales and scattered fine erect setae; setae primarily arranged along inner margin of femora and tibiae. Femora clavate, widest at apical ⅓, simple, lacking tooth. All tibiae more or less straight in female, slightly inwardly arcuate in male; inner margin with small to minute rounded asperites in apical ½ in female, with numerous large teeth throughout length in male; mucro large, curved, smaller on meso- and metatibia. Metatibia with apical bevel broad, glabrous; api- cal comb of setae not ascended along outer margin of tibia; mucro small, apical margin of metatibia excised adjacent to base of mucro such that a second tooth is evident on the margin at the apex of the excision. Tarsal groove squamose (scales may be abraded), with only one or two large appressed scales. Tarsi elongate, article 1 slightly longer than article 2, article 2 slightly longer than article 3; article 3 bilobed, article 4 elongate, extended beyond apices of 3 by slightly greater than length of article 3; ventrally with elongate, moderately dense, fine, wispy vestiture on articles 1 and 2, very dense and pilose on article 3; claws simple, divergent, free.

Prosternum with procoxae contiguous, situated slightly closer to anterior margin than to posterior margin. Mesocoxae proximate, separated by more or less ¼ diameter of mesocoxa. Mesosternum sparsely punctate, lacking scales. Mesepimeron short-trapezoidal, anterior margin directed to elytra then turned anteriorly so that mesepisternum contacts elytron in a strip near extreme base of elytron. Metasternum short, concave medi- ally; round scales present only laterally. Metepisternal suture present, distinct; metepisternum extremely narrow, about 10 times as long as wide. Metacoxae widely separated by about diameter of a metacoxa.

Abdomen with sparse, rounded scales laterally and very sparse, very fine erect setae throughout. Visible sternite 1 subequal in length to 2; 3 and 4 subequal in length, short, their combined length slightly shorter than length of visible sternite 2; visible sternite 5 longer than length of 3 and 4 combined. Base of visible sternite 1 concave in male, flat to slightly convex in female. Apex of visible sternite 5 rounded in male and female. Tergite 7 transverse in both male and female, with apical margin shallowly medially emarginate or not; posterolateral angles rounded, not projected.

Male genitalia. Sternite 8 large, trapezoidal, posterior margin moderately emarginate and deeply cleft; sternite 9 long, broad. Tegmen well sclerotized, parameres developed, directed anteriad, very lightly sclerotized. Aedeagus cylindrical, elongate, sclerotized throughout; apex produced, slightly reflexed, no apical setae present. Apodemes slightly shorter than length of aedeagus.

Female genitalia. Sternite 8 small, flat, subrhomboidal, as wide as long, widest at base; very lightly sclerotized; apical ⅓ with elongate, erect setae; apodeme very narrow, from 4 to 8 times length sternite. Ovipositor (very pale and indistinct in some specimens) ¼ length abdomen or less, setae present, short, sparse, apical; baculi absent; hemisternites (where apparent) short or long, subequal in length, not apparently subdivided into basal and apical portions; in dorsal view fused at apex; styli absent. Spermatheca subcylindrical, point of insertion of duct not developed, point of insertion of gland globose, proximal.

KEY TO SPECIES OF COCONOTUS

1 Pronotum widest at or near base (figs. 11, 13). Elytra flattened and broadly oval in dorsal view, especially so in males (figs. 11, 13). Female with humeral carina short, distinct, and sharp................................. C. williamsi Anderson and Lanteri

1′ Pronotum widest from anterior ⅓ to middle (figs. 24, 26, 36). Elytra more convex and elongate-oval in dorsal view (figs. 24, 26, 36). Female with humeral carina short, low, rounded or short, very distinct, laterally produced, acute....................... 2

2 Female with humeral carina short, low, round- ed (fig. 24). Male with subapical callus lacking.... C. kuscheli Anderson and Lanteri

2′ Female with humeral carina short, very distinct, laterally produced, acute (fig. 36). Male not known........................ C. tuberculatus Anderson and Lanteri

Coconotus williamsi Anderson and Lanteri , new species Figures 10–22

TYPES: Holotype male labeled ‘‘ COSTA RICA. Prov. Puntarenas./ P . N . Isla del Coco. Bahía Wafer. / 1 m. Oct.1994. J . F . Quesada. / Long:-87:03:30 Lat:5:32:45 #3314,’’ with INBio barcode label 2544747 ( INBio). Allotype female labeled ‘‘ Bahía Chatan , P . N . Isla del Coco ,/ Prov. Punt., COSTA RICA. 5 a 9/feb 1993. F . Quesada, L-S-0 0,’’ with IN- Bio barcode label 1851622 ( INBio). Paratypes as follows: Costa Rica. Cocos Island . 3– 13.IX.1905, F . X. Williams (1♀; CASC). 8.III.1964, G . Kuschel (23; NZAC). Bahía Chatan , 5–9.II.1993, F . Quesada (43, 3♀; INBio, CMNC, MZLP; 1851918, 1851621, 1366654, 1851623, 1851624, 1851626, 1851625, 1851627). Bahía Chatan , 5– 9.II.1993, P . Rios (1♀; CMNC; 1850791). Bahía Wafer , 13, X.1994, J . F . Quesada (13, 1♀; INBio, CMNC; 2544771, 2544746). Bahia Yglesias a la Catarata, 20 m, 21.XII.1997, C . Flores, E . Ulate (43, 1♀, AMNH, CMNC, INBio ; 3033085, 3033086, 3033087, 3033088, 3033089, Cuesta el Gallinero, sendero a Cerro Yglesias, 200 m, 28.XII.1997, 19.XII.1997, C . Flores, E . Ulate (2♀; AMNH, INBio; 3033002, 3033151). El Guarumal, sendero Wafer a Chatan , 30 m, C . Flores, E . Ulate (13; INBio; 3033022). Los Llanos , 260 m, 24.XII.1997, C . Flores, E . Ulate (2 3; INBio; 3033066, 3033067). Orilla del Rio Genio , 10 m, 17.XII.1997, C . Flores, E . Ulate (23, 2♀; CMNC; 3033075, 3033076, 3033078, 3033079). Sendero Las Cuevas , 20 m, 16.XII.1997, C . Flores, E . Ulate (13, 1♀; INBio; 3033013, 3033014). Total paratypes, 13♀, 183 .

ETYMOLOGY: This species is named after Francis X. Williams of the California Academy of Sciences, who collected the first known specimen during fieldwork on Cocos Island (and the Galapagos) in 1905–6.

DIAGNOSIS: This species is easily distinguished by the form of the pronotum, which is widest at the base, the flattened and broadly oval elytra, and the presence of metallic blue or green scales.

DESCRIPTION: Male, length 7.8–8.1 mm, width 3.8–4.0 mm. Female, length 6.1–6.6 mm, width 2.8–3.2 mm. Scales green, blue, or gold, with metallic reflection (greasy and appearing black in some specimens); dense along lateral margins pronotal disk and pronotal flanks, entire elytra, and laterally on metasternum and metepisternum, otherwise very sparse or absent; not forming distinct elytral pattern. Rostrum regularly, finely punctate in basal ⅔; very sparsely and finely punctate in apical ⅓; distinctly deflexed api- cally. Frons with shallow but distinct fovea between eyes. Pronotum widest at or near base; disk finely indistinctly punctate, punctures larger and more irregular laterally. Elytra widest at or near midlength. Elytral disk moderately inflated in female; very flat, broadly flattened with distinct deflexed lateral margins in male. Elytral punctures confused, distinct intervals not evident except medially on disk; intervals uniformly flat; base of interval 7 with short, distinct, sharp humeral carina in female. Subapical callus present, low, at apex of deflexed lateral margin. Apical declivity with erect setae short, indistinct. Visible sternite 5 of female flat; lateral margins more or less convergent from base to apex.

Male genitalia. Aedeagus in lateral view moderately curved; in dorsal view with apex moderately produced, tip sharply acuminate; apical ⅓ (at median orifice) and basal regions expanded, distinctly wider than intervening length and apical region. Internal sac visible at base of aedeagus, with no visible internal sclerotization.

Female genitalia. Tergite 7 with posterior margin rounded. Sternite 8 with apodemes about four times length sternite. Ovipositor with hemisternites short, about ½ total length of sternite 8 (including apodemes).

DISTRIBUTION: COSTA RICA. Cocos Island. Bahía Chatan, Bahía Wafer, Bahia Yglesias, Cuesta el Gallinero, El Guarumal, Los Llanos, Orilla del Rio Genio, Sendero Las Cuevas.

NATURAL HISTORY: As far as known, this species is native and endemic to Cocos Island. No details are known of habitat association or of food habits. Specimens have been collected at or very near sea level, apparently along the coast (‘‘bahía’’ = bay).

Coconotus kuscheli Anderson and Lanteri , new species Figures 23–35

TYPES: Holotype male labeled ‘‘ Bahía Chatan , P . N . Isla del Coco ,/ Prov. Punt., COS- TA RICA. 5 a 9/feb 1993. F . Quesada, L-S- 0 0’’, with INBio barcode label 1851721 (IN- Bio). Allotype female labeled as holotype, with INBio barcode label 1851628 ( INBio). Paratypes as follows: Costa Rica. Cocos Island . 8–9.III.1964, G . Kuschel (33, 2♀; CMNC, NZAC). Bahía Chatan , 5–9.II.1993, F . A . Quesada (33; INBio, CMNC, MZLP; 1851722, 1851720, 1851913). Bahía Wafer , 13, X.1994, J . F . Quesada (1♀; CMNC; 2544744). Los Llanos , 260 m, 24.XII.1997, C . Flores, E . Ulate (13, 1♀; CMNC, INBio; 3033059, 3033060). Total paratypes, 4♀, 73 .

ETYMOLOGY: This species is named after Guillermo (Willy) Kuschel of New Zealand, who collected the first known specimens of this species while conducting field work on Cocos Island (and the Galapagos) in 1964.

DIAGNOSIS: This species can be recognized by the form of the pronotum, which is widest from the midlength to the anterior ⅓, by males lacking a subapical callus, and by females with the humeral carina short, low, and rounded.

DESCRIPTION: Body length male 5.5–5.7 mm, width 2.3–2.4 mm. Body length female 6.2–6.8 mm, width 2.8–3.0 mm. Scales tan, dark brown, or golden, with indistinct metallic reflection (greasy and appearing black in some specimens); dense on pronotal disk and flanks, elytra, metasternum, laterally on abdominal sternites 1 and 2 and sternite 5, otherwise very sparse or absent; not forming distinct elytral pattern. Rostrum regularly, finely punctate in basal ⅔; very sparsely and finely punctate in apical ⅓; very slightly deflexed at apex. Frons with short, moderately deep basal sulcus terminated basally in shallow fovea. Pronotum widest at or near midlength; disk distinctly, irregularly punctate. Elytra widest at or behind midlength. Elytral disk not distinctly inflated or flattened. Elytral punctures regularly, serially arranged; intervals distinct throughout; intervals flat (sutural interval 6) or uniformly convex (intervals 7–10), except extreme bases of intervals 3 and 5 that are slightly swollen in male,

Female genitalia. Tergite 7 with posterior margin emarginate at middle. Sternite 8 with apodemes about eight times length sternite. Ovipositor with hemisternites short, about ⅓ total length of sternite 8 (including apodemes).

DISTRIBUTION: COSTA RICA. Cocos Island. Bahía Chatan, Bahía Wafer, Los Llanos.

NATURAL HISTORY: As for C. williamsi .

Figures

Figs. 1–2. Galapagonotus cuneiformis (Waterhouse), female. 1, Lateral habitus; 2, dorsal habitus.

Fig. 3–9. Galapagonotus cuneiformis (Waterhouse). 3, Male abdomen, ventral view; 4, female abdomen, ventral view; 5, ovipositor, sternite 8, tergites 7 and 8; 6, spermatheca; 7, aedeagus, lateral view; 8, aedeagus, dorsal view; 9, sternites 8 and 9, tergite 8. Scale line = 1 mm, except spermatheca, = 0.25 mm.

Figs. 10–13. Coconotus williamsi Anderson and Lanteri. 10, Lateral habitus, female; 11, dorsal habitus, female; 12, lateral habitus, male; 13, dorsal habitus, male.

Figs. 14–22. Coconotus williamsi Anderson and Lanteri. 14, Male abdomen, ventral view; 15, female abdomen, ventral view; 16, sternite 8; 17, tergites 7 and 8; 18, hemisternites; 19,spermatheca; 20, aedeagus, lateral view; 21, aedeagus, dorsal view; 22, sternites 8 and 9, tergite 8. Scale lines = 1 mm, except spermatheca = 0.25 mm.

Figs. 23–26. Coconotus kuscheli Anderson and Lanteri. 23, Lateral habitus, female; 24, dorsal habitus, female; 25, lateral habitus, male; 26, dorsal habitus, male.

Figs. 27–35. Coconotus kuscheli Anderson and Lanteri. 27, Male abdomen, ventral view; 28, female abdomen, ventral view; 29, sternite 8; 30, tergites 7 and 8; 31, hemisternites; 32, spermatheca; 33, aedeagus, lateral view; 34, aedeagus, dorsal view (inset shows large curved sclerite of internal sac at base of aedeagus); 35, sternites 8 and 9, tergite 8. Scale line = 1 mm, except spermatheca = 0.25 mm.

Fig. 36. Coconotus tuberculatus Anderson and Lanteri, dorsal habitus, female.

Figs. 37–41. Coconotus tuberculatus Anderson and Lanteri. 37, female abdomen, ventral view; 38, sternite 8; 39, tergites 7 and 8; 40, hemisternites, 41, spermatheca. Scale line = 1 mm, except spermatheca = 0.25 mm.