Heterocerus gnatho, LeConte, 1863

King, Jonas G. & Lago, Paul K., 2012, The variegated mud-loving beetles (Coleoptera: Heteroceridae) of Mississippi and Alabama, with discussion and keys to the species occurring in the southeastern United States, Insecta Mundi 2012 (275), pp. 1-53 : 10

— Elytra and pronotal coloration varying from entirely dark brown to yellow with pale pronotal margins ( Fig. 14 ); hypermandibulate males common; genitalia with posterior edge of the dorsal plate more deeply notched ( Fig. 37 ); medial edge of parameres diverging slightly ( Fig. 37 ) ..... ..................................................................................................................... 2. H. gnatho LeConte

Figures

Figures 11-17. Habiti of large Heterocerus species lacking post-mesocoxal lines. 11) H. pallidus (male is hypermandibulate). 12) H.intermuralis (male is hypermandibulate). 13) H.fatuus. 14) H. gnatho. 15) H.longilobulus. 16) H. sandersoni. 17) H. angustatus.Scale line = 1mm.

Figures 32-44. Male genitalia of southeastern Heterocerus species. The ventral view is shown unless otherwise stated. 32-39) Species that lack post-mesocoxal lines. 32) H. fatuus. 33) H. glicki. 34) H. intermuralis. 35) H. angustatus; left - ventral view, right - dorsal view. 36) H. longilobulus. 37) H. pallidus; left - ventral view, right - dorsal view. 38) H. gnatho; left - ventral view, right - dorsal view. 39) H. sandersoni; left - ventral view, center - dorsal view, left - lateral view. 40-44) Species with post-mesocoxal lines. 40) H. sinuosus; left - dorsal view, right - ventral aspect. 41) H. mollinus. 42) H. insolens; left - dorsal view, right - ventral aspect. 43) H. parrotus. 44) H. brunneus.