Coenosiini Verrall, 1888

Michelsen, Verner, 2022, Costal vein chaetotaxy, a neglected character source in Fanniidae and Muscidae (Diptera: Calyptratae), European Journal of Taxonomy 826, pp. 94-134 : 124-127

Tribe Coenosiini Verrall, 1888

A well defined, conceivably monophyletic tribe ( Couri & Pont 2000), currently with 34 recognized genera; species from 23 genera were examined ( Table 1 ).

• Anaphalantus Loew, 1857 , Apsil Malloch, 1929 , Bithoracochaeta Stein, 1911 , Brevicosta Malloch, 1921 , Cephalispa Malloch, 1935 , Limnospila Schnabl, 1902 , Lispocephala Pokorny, 1893 , Macrorchis Rondani, 1877 , Neodexiopsis Malloch, 1920 , Notoschoenomyza Malloch, 1934 , Orchisia Rondani, 1877 , Oxytonocera Stein, 1919 , Parvisquama Malloch, 1935 , Pseudocoenosia Stein, 1916 , Pygophora Schiner, 1868 , Reynoldsia Malloch, 1934 , Schoenomyza Haliday, 1833 , Schoenomyzina Malloch,1934 , Spanochaeta Stein, 1919 , Spathipheromyia Bigot, 1884 , Stomopogon Malloch, 1930

All examined species from these 21 genera show the prevalent muscid state A 4 in which vein C is bare dorsally and extensively setulose ventrally: Anaphalantus longicornis (Macquart, 1843) , Apsil (7 spp), Bithoracochaeta (4 spp), Brevicosta (1 sp. indet.), Cephalispa (1 sp. indet.), Limnospila albifrons (Zetterstedt, 1849) , Lispocephala (10 spp), Macrorchis meditata (Fallén, 1825) , Macrorchis sp. indet., Neodexiopsis (3 sp. indet.), Notoschoenomyza sulfuriceps Malloch, 1934 , Notoschoenomyza sp. indet., Orchisia costata (Meigen, 1826) , Oxytonocera nigrohalterata (Stein, 1904) , Parvisquama (3 spp indet.), Pseudocoenosia abnormis Stein, 1916 , P. solitaria (Zetterstedt, 1838) , Pygophora (7 spp), Reynoldsia (3 spp), Schoenomyza (3 spp), Schoenomyzina triangularis Malloch, 1934 , Schoenomyzina sp. indet., Spanochaeta dorsalis (Roser, 1840) , Spathipheromyia atra Malloch, 1934 , S. guttipennis (Thomson, 1869) , Stomopogon albiseta (Stein, 1911) and S. inculta (Stein, 1911) .

The examined species of Neodexiopsis , Orchisia and Spathipheromyia have the setulae from the lower anterior spinule-setula row displaced dorsally, so that they occupy the usually bare strip separating the spinule-setula rows.

• Coenosia Meigen, 1826 ( Figs 1A–B , 18A–D )

The examined 50+ species of Coenosia all show the prevalent muscid state A 4 in which vein C is bare dorsally and extensively setulose ventrally ( Figs 1A–B , 18A–C ). One exception was found in the boreal Palearctic species Coenosia emiliae Lukasheva, 1986 ( Fig. 18D ), which has dorsal costal setulae occupying CS2 and CS3 (state A7).

• Cordiluroides Albuquerque, 1954

The examined species, Cordiluroides bistriata (Wulp, 1896) and Cordiluroides sp. indet., agree with Coenosia emiliae in having setulae dorsally on CS2 and CS 3 in addition to the mandatory ventral costal setulae (state A7).

Figures

Fig. 1. Coenosia pudurosa Collin, 1953, ♀. A. Detail of wing, ventral view, showing different kinds of macrotrichia: cs = campaniform sensillum; gs = gustatory sensillum; se = setula; sp = spinule. B. Wing, ventral view showing costal sectors 1‒4: CS = costal sector.

Fig. 18. ♀♀ of Coenosia Meigen, 1826. A. C. intermedia (Fallén, 1825), wing, ventral view. B. Same, detail of wing showing CS1‒3, ventral view. C. Same, detail of wing showing CS1‒3, dorsal view. D. C. emiliae Lukasheva, 1986, detail of wing showing CS1‒3, dorsal view.