Honca Pogue, 2019

Brown, John W., 2019, New genera, new species, and new combinations in New World Cochylina (Lepidoptera: Tortricidae: Tortricinae), Zootaxa 4671 (2), pp. 195-222 : 203-204

Honca Pogue , new genus

Type species: Phalonia grandis Busck, 1907 , by monotypy.

Honca Pogue and Mickevich, 1990: 322 (nomen nudum).

Cochylini New Genus 4 [ grandis group]: Brown 2005: 208; Metzler and Brown 2014: 278.

Honca is described to accommodate the single species Phalonia grandis , which is among the largest Cochylina in the Nearctic region. The forewing pattern ( Fig. 7 ) somewhat resembles that of Henricus umbrabasanus , however, H. grandis is much larger (forewing length 11‒13 mm vs. 8‒10 mm) and lacks a pale brown basal patch. Honca also resembles Aphalonia Razowski, 1984 , a South American genus, in size and forewing maculation, however, the genitalia of the two have little in common.

Diagnosis. The shapes of the valva and transtilla of Honca are similar to those of some species of Aethes , however, Honca lacks the sickle-shaped socii that are characteristic of most species of Aethes . The most conspicuous autapomorphy of Honca is the highly modified phallus-juxta complex in the male genitalia ( Fig. 23 ): the juxta is rather long, slender, subrectangular, and widely bifurcate distally, and is strongly attached to the large, curved phallus. The phallus has a long, flattened process of the right side that is as long as the phallus, and a short, stout, more rounded process on the left side that is about one-fourth the length of the phallus. The latter features are reminiscent of the paired process that subtend the phallus in some species of Aethes . In the female genitalia the spiculate, clubshaped arms of the sterigma ( Fig. 37 ) are somewhat reminiscent of some species of Eupinivora . Honca is further characterized by the absence of ocelli (usually present in Cochylina), a 3-segmented maxillary palpus (shared with Eugnosta ), a reduced haustellum (shared with Eugnosta ), four acanthi in the female frenulum (occurring consistently nowhere else in Cochylina), fusion of the socii with the tegumen, and the absence of cornuti in the vesica (shared with Cochylis , Thyraylia , and Cochylichroa, and few other genera).

Description. Head: Vertex and upper frons rough scaled, lower frons with sparse appressed scales; ocellus absent; sensory setae of antenna about as long as flagellomere diameter; labial palpus porrect, combined length of all segments ca. 3 times horizontal diameter of compound eye, shorter in female, segment III nearly concealed by scaling of segment II; maxillary palpus 3-segmented. Thorax: Posterior crest absent. Forewing length 11.0– 12.5 mm in male, 11.5–13.0 mm in female; length 2.4–2.8 times maximum width; forewing with base of costa straight, gently curved in distal 0.33 to rounded apex, costal fold absent; termen straight, oblique; Sc less than 0.5 wing length; R 1 originating beyond middle of discal cell; R 2 nearer to R 3 than to R 1; R 5 terminating at costa; M 3 and CuA 1 separate; CuA 2 originating ca. 0.8 length of discal cell; CuP absent; A1+2 stalked about 0.5 of their lengths. Hindwing with costa mostly straight, gently arched in apical 0.33, male without costal roll, termen slightly concave beneath apex; Sc+R 1 ca. 0.8 wing length; Rs and M 1 stalked to ca. 0.6 length; M 3 and CuA 1 separate; CuA 2 originating ca. 0.66 length of discal cell; frenulum with one acanthus in male, four in female. Abdomen: Unmodified. Male genitalia ( Fig. 23 ) with uncus absent; socius prominent, fused to tegumen; transtilla with large, triangular, median process, with strongly sclerotized, pointed tip; valva short, broad, with upturned, well-sclerotized costa and attenuate apex, sacculus a straight sclerotized margin along venter of valva with slightly produced distal tip; vinculum arms con- nected distally by membrane; juxta a complicated plate, narrow, subrectangular, deeply emarginate on distal 0.35 distally resulting in pair of pointed processes, basal end fused to phallus. Phallus large, ca. 0.8 times length of valva, subtended by long, flattened lateral process on right side and shorter, peglike process on left (possibly homologous with “anellus lobes” of Sabourin et al. 2002), coecum large; vesica without cornuti. Female genitalia ( Fig. 37 ) with papillae anales weakly curved, sclerotized; apophyses short, stout, apophyses anteriores ca. 1.1 times length of apophyses posteriores; sterigma membranous, median area with a pair of mesially-directed, spiculate, slightly club-shaped processes originating from posterior base of apophyses anteriores, each with 15‒20 long setae; ostium bursae wide; colliculum absent; ductus bursae short, undifferentiated from corpus bursae; corpus bursae with vertical lines of sclerotization below ostium bursae and with large patch of minute spicules; frail accessory bursae originating anterad of middle of ductus bursae.

Etymology. The generic name is an arbitrary combination of letters and is feminine in gender.

Figures

FIGURES 1–10. Adults of Cochylini genera. 1. Anacochylidia maderana, Madera Canyon, Santa Rita Mountains, Arizona (EME). 2. Cagiva cephalanthana, Dorman Lake, Oktibbeha Co., Mississippi (USNM). 3. Atroposia oenotherana, Houston, Harris Co., Texas (USNM). 4. Atroposia oenotherana, Radisson, Onondaga Co., New York (USNM). 5. Cybilla hubbardana, Shafter, Presidio Co., Texas (USNM). 6. Cybilla hubbardana, Lordsburg, Hidalgo Co., New Mexico (CNC). 7. Honca grandis, Chimney Gulch, Golden, Colorado (USNM). 8. Pogospinia floridana, Long Key State Park, Monroe Co., Florida (USNM). 9. Nycthia pimama, Portal, Cochise Co., Arizona (USNM). 10. Nycthia yuccatana, Falcon State Park, Starr Co., Texas (USNM).

FIGURES 19–26. Male genitalia of Cochylini genera. 19. Anacochylidia maderana, Palmerlee, Arizona, USNM slide 23,359. 20. Atroposia oenotherana, Padre Island National Seashore, Texas, USNM slide 153,596. 21. Cagiva cephalanthana, Dorman Lake, Mississippi, USNM slide 150,530. 22. Cybilla hubbardana, Tucson, Arizona, USNM slide 29,397. 23. Honca grandis, Ogden Canyon, Utah, USNM slide 153,598. 24. Nycthia pimama, Blair Valley, California, USNM slide 144,042. 25. Nycthia yuccatana, Laguna Atacosa, Texas, USNM slide 120,405. 26. Pogospinia floridana, Key Largo, Florida, USNM slide 23,249.

FIGURES 33–38. Female genitalia of Cochylini genera. 33. Anacochylidia maderana, Palmerlee,Arizona, USNM slide 23,358. 34. Atroposia oenotherana, Padre Island National Seashore, Texas, USNM slide 153,597. 35. Cagiva cephalanthana, Dorman, Mississippi, USNM slide 153,609. 36. Cybilla hubbardana (holotype), Tucson, Arizona, USNM slide 23833. 37. Honca grandis, Wellsville, Utah, USNM slide 153,599. 38. Nycthia pimama, Miller Canyon, Arizona, slide MGP 660.