Argentinolycus, Matallanas, Jesús & Corbella, Cecília, 2012

Matallanas, Jesús & Corbella, Cecília, 2012, Redescription of Iluocoetes Jenyns, 1842; proposal of a new genus, Argentinolycus, for Iluocoetes elongatus (Smitt, 1898), and description of Patagolycus melastomus gen. et sp. nov. (Teleostei, Zoarcidae), Zootaxa 3296, pp. 1-18 : 6-7

Argentinolycus gen. nov.

( Fig. 10 ; Tables 2−3)

Type species: Phucocoetes variegatus elongatus Smitt, 1898 .

A synonymy is found in Gosztonti (1977). Owing to misidentifications ( Regan, 1913; Norman, 1937), the first available scientific name in print for this form is Smitt’s (1898: 43) Phucocoetes variegatus effusus . As this name was not used until Anderson and Gosztonyi (1991: 2 in key only), the prevailing usage of elongatus is used here for nomenclatural stability.

The diagnosis given below is taken from the diagnosis and description of Iluocoetes given by Anderson (1994: 61−63), based mainly on a 147 mm SL specimen of Iluocoetes elongatus ( CAS 53295), and also from the description of Iluocoetes elongatus by Gosztonyi (1977). Skeletal structures are represented by Anderson (1994): cranium (8, fig. 3D: dorsal view of neurocranium; 62, fig. 112: left lateral view of neurocranium; fig. 113: bones of left side of head showing suborbital configuration; fig. 114: left splachnocranium and opercular bones), pectoral girdle (62, fig.115: left pectoral girdle) and caudal skeleton (63, fig. 116: two specimens, 147 and 131 mm SL, both CAS 53297).

Diagnosis. Neurocranium elongate, depressed; parasphenoid wing broad, but without dorsal ramus projecting above ventral base of trigeminofacialis foramen; frontal and parasphenoid articulating; pterosphenoid enlarged; intercalar very small, set posteriorly. Frontal bones separate; frontal corner tapering; parietals meeting in dorsal mid-line; supraoccipital small; supraoccipital and exoccipital narrowly articulating posteriorly; sphenotic excluded from parietal by frontal and pterotic.

Posterior ramus of hyomandibula elongate; palatopterygoid series well developed; ectopterygoid overlap both anterior and dorsal surface of quadrate. Ceratohyal–epihyal juncture with bone interdigitating along its entire length; branchiostegal rays 6. Suborbital bones 7–8, canal with 6 pores. Posttemporal ventral ramus absent; scapular foramen enclosed, scapula with well developed posterior strut; postcleithrum present. Vertebrae asymmetrical, 22–24+62–69 = 84–90.

Oral valve well developed; gill slit extending ventrally to slighly below lower end of pectoral-fin base; vertebrae asymmetrical; no interorbital or occipital pores and no commissure across parietals; 6 suborbital pores along ventral ramus (6+0); two nasal pores; only postorbital pore 4; lateral line mediolateral complete; pyloric caeca absent; pelvic-fin membranes excised at tip; scales, palatine and vomerine teeth present. The character state of other diagnostic features of this genus can be observed in Tables 2 & 3.

Description. A complete description of Argentinolycus elongatus ( Smitt, 1898) , type species of the new genus, can be found under Iluocoetes elongatus ( Smitt, 1898) in Gosztonyi (1977: 215–217); other data are in Gosztonyi (1984, 1988).

Etymology. From Argentina, as the species is known mainly from the Argentine Patagonia, and the Greek lykos (wolf), a commonly used suffix for southern hemisphere zoarcid genera. Gender: masculine.

Figures

FIGURE 10. Interrelationships among the Magellanic endemic lycodine genera. Consensus tree resulting from the parsimony analysis of 78 morphological characters (Length = 105; CI = 0.53; RI = 0.50). Bootstrap support values (rep = 1000) are indicated on the nodes. White boxes on branches: homoplastic apomorphies; black boxes: synapomorphies.