Calciphilopteris wallichii (J.Sm.) Yesilyurt & H.Schneider

Yesilyurt, Jovita C. & Schneider, Harald, 2010, The new fern genus Calciphilopteris (Pteridaceae), Phytotaxa 7, pp. 52-59 : 57

4. Calciphilopteris wallichii (J.Sm.) Yesilyurt & H.Schneider , comb. nov. — Fig. 1C .

Basionym:— Doryopteris wallichii Smith (1841: 404) , Lithobrochia smithii T. Moore (1862: 342) , nomen nudum, based on D. wallichii J.Sm. Holotype: PHILIPPINES: Luzon, Cuming 238 (K, isotypes BM, C, G, L).

Rhizome usually short-creeping, scales 2.5(–3) mm, light brown, lanceolate to long-lanceolate, margin entire to rarely dentate. Petiole terete to slightly sulcate towards the base of the lamina; dark brown, atropurpureous, with fibrils. Sterile and fertile fronds sub-coriaceous; glabrescent, indumentum on the abaxial side with glandular hairs and fibrils on the main veins. Sterile fronds up to 30 cm; lamina up to 17 cm, ranging from simple, entire to sagittate, hastate, or basally lobed; hydathodes generally present on the upper surface only (sometimes obscure). Fertile fronds up to 40 cm; lamina up to 19 cm; sagittate, hastate, sub-pedate to rarely pedate, with 2(–3) pairs of pinnae/segments; basal pinna/segment falcate to lanceolate, conspicuously ascendant, with 1–2 basiscopic segment/lobes, which are ascendant to usually parallel with the petiole; other pinnae/segment broadly lanceolate to lanceolate, usually patente. Indusium usually with entire margin. Sporangium with pedicel 1–1½ × the capsule length; capsule with annulus with 20 (24) indurated cells. Spores usually castaneous to light castaneous, cristate.

Distribution:— Philippines (mainly Luzon).

Selected specimens:— PHILIPPINES: Copeland 95 (BM, C, S); Curran 15608 (BM, C). Elmer 6349 (C. NY); Price 2007 (BM, L, US); Ramos 33245 (BM, L); Topping 857 (C, GH, MO, NY, S); 1014 (GH, NY); 1247 (GH, US); 1219 (GH, NY); Williams 1501 (PE, UC).

Figures

FIGURE 1. Sketches of fertile leaves showing their general attributes and morphological variation: A. Calciphilopteris ludens. B. Calciphilopteris papuana. C. Calciphilopteris wallichii.