Deltochilum (Aganhyboma) larseni, Silva, Fernando A. B., Louzada, Júlio & Vaz-De-Mello, Fernando, 2015

Silva, Fernando A. B., Louzada, Júlio & Vaz-De-Mello, Fernando, 2015, A revision of the Deltochilum subgenus Aganhyboma Kolbe, 1893 (Coleoptera: Scarabaeidae: Scarabaeinae), Zootaxa 3925 (4), pp. 451-504 : 478

Deltochilum (Aganhyboma) larseni new species

( Figs 14 , 91 , 113 )

Etymology. A patronym honoring Trond Larsen, great and enthousiatic scarab ecologist who collected most of the type specimens.

Description. Body. Black or dark brown coloration, with obvious sheen ( Fig. 113 ). Length <13 mm. Head. Punctures dense, nearly touching; around clypeus shallower, smaller and more dispersed than those on the interocular surface. Lateral margin of gena, around eyes, without row of setae or with almost inconspicuous setae. Dorsal interocular distance approximately 2.5 times eye width. Thorax. Surface of pronotum with dense foveiform punctures. Punctures on the sides, anterior and posterior angles nearly touching. Discal punctures smaller and more dispersed than those of lateral portion, separated approximately by one diameter. Anteromedial punctures smaller and more dispersed than discal punctures. Posteromedial punctures usually separated by at least one diameter. Margin between anterior and lateral angles with row of small setae. Margin between lateral and posterior angles with length somewhat shorter than that of margin between anterior and lateral angles. Hypomera with dense foveiform punctures on anterior portion and dispersed on posteromedial portion; anteromedial punctures almost touching. Diagonal band of hypomera from posteromedial portion to middle of lateral margin with dispersed punctures on otherwise smooth surface. Elytra. Lateral margins slightly curved outward ( Fig. 113 ) and surface slightly irregular (rugose). Elytral striae with carinate margins clearly separated; strial punctures separated by regular spacing; most of punctures round intermingled with some slightly elliptical. Interstriae flattened, with foveiform punctures surrounded by approximately five obvious shiny microtubercles. Foveiform punctures separated on elytral disc by approximately one diameter. Basal carina of seventh interstria elongate, length about four times the width of the seventh interestria. Basal carina of ninth interstria with approximately one-half of elytral length (as in Fig. 80). Apical carina or tubercle of second elytral interstria absent. Apical carina of third interstria weaker than apical carina of fourth interstria. Length of apical carina of sixth interstria longer than that of apical carina of fifth interstria. Length of apical carina of seventh interstria longer than or equal to the sum of lengths of apical carinae of fifth and sixth interstriae. Pseudepipleural carina incomplete, effaced medially (as in Fig. 80).

Material studied. Holotype: ECUADOR: SUCUMBIOS: R.P.F. Cuyabeno, Trocha Zábalo-Güepí, km 10, 9.VIII.2010, Colección Manual nocturna, Bosque de tierra firme colinado, Pablo Araujo, LOTE 557 (1 ♂ CEMT).

Paratypes [15]: ECUADOR: SUCUMBIOS: R.P.F. Cuyabeno, Trocha Zábalo-Güepí, km 10, 9.VIII.2010, Colección Manual nocturna, Bosque de tierra firme colinado, Pablo Araujo, LOTE 557 (1 ♂ CEMT); PASTAZA: B.P. Oglán, T4, Bsvpm. 554 msnm, 201146 E 9853376N, 23.VIII.2008, Col. W. Chamorro & V. Guasumba (1 ♀ CEMT). PERU: valle cerca Abancay, 2.VII.1948, C.S. Carbonell (1 ♂ MZUSP); Upper Rio Marañón, 8.X.1928 (1 ♂ MZUSP); MADRE DE DIOS: Río Tambopata, 300 m, Ccolpa de Guacamayos, 13°08.5'S, 69°36.4'W, A Forsyth, X.1995 (1 ♂ AFIC); Rio Madre de Dios, Rio Los Amigos, base camp, 1° terra firme, 290 m, pitfall crushed but live spirobolid millipede, 12°34'10.0"S, 70°06'01.4"W, T. Larsen, IV.2000 (1 ♂ CEMT, 1♂ 1♀ AFIC); same but 12–13.IV.2000 (1♂ CEMT); same but Rio Amiguillos, small rever flood plain, 260 m, pitfall millipede, 12°22'25.4"S, 70°22'13.2"W, T. Larsen, V.2000 (1♂ AFIC, 1♀ CEMT); same but Rio Palma Real Grande, Limon Camp, 12°32'20"S, 68°51'40"W, Flight intercept trap, 220 m, T. Larsen, 11–12.X.1999 (1♀ CEMT); same but pitf. crished millipede, 12–13.X.1999 (1♀ AFIC); same but perching night, 11.X.1999 (1♀ AFIC); same but 12°32'20"S, 68°51'41"W, Flight interception trap, 400 m, 2–3.IV.1999 (1♀ AFIC).

Habitat. Ecuador and Peru ( Fig. 91 ), from available data this species inhabits Amazon rainforest.

Diagnosis and remarks (within the acropyge complex): length <13 mm; punctures of pronotal disc separated by approximately one diameter; posteromedial punctures of pronotum usually separated by at least one diameter; pronotal margin, between lateral and posterior angles, with length somewhat shorter than margin between anterior and lateral angles; elytral striae with carinate margins clearly separated; strial punctures separated by regular spacing; most of strial punctures round intermingled with some slightly elliptical; apical carina or tubercle of second interstria absent; pseudepipleural carina incomplete, effaced medially (as in Fig. 80). The following aedeagus characters are also diagnostic: parameres symmetric and short, length shorter than phallobase; in lateral view forming an angle of approximately 45° with phallobase ( Fig. 14 ). Lamella copulatrix (LC): subrectangular sclerites. Fronto-lateral peripheral sclerite (FLP) as in Fig. 54.

Figures

FIGURES 12 – 19. Aedeagus in Deltochilum (Aganhyboma). 12. D. schefflerorum new species. 13. D. acropyge new status. 14. D. larseni new species. 15. D. acanthus. 16. D. longiceps new status. 17. D. valgum. 18. D. icaroides. 19. D. kolleri new species. Figures scale = 1 mm.

FIGURES 89 – 92. Known distribution of D. (Aganhyboma). 89. trisignatum complex. 90. cupreicolle complex. 91. acropyge complex. 92. finestriatum complex.

FIGURES 111 – 118. Species of D. (Aganhyboma). 111. D. acanthus. 112. D. arturoi new species. 113. D. larseni new species. 114. D. cangalha new species. 115. D. alpercata new species. 116. D. finestriatum new species. 117. D. icaroides. 118. D. valgum. Figures scale = 1 mm.