Anisoscelini, Latreille, 1829

Leavengood, John M., Brailovksy, Harry & Barrera, Ernesto, 2024, Studies in the New World tribe Anisoscelini Laporte, 1832 (Heteroptera: Coreidae: Coreinae): updated distributions, nomenclatural changes, and keys to the genera of the tribe and the species of Anisoscelis Latreille, 1829 and Bitta Osuna, 1984, Zootaxa 5415 (2), pp. 241-268 : 262-264

publication ID

https://doi.org/ 10.11646/zootaxa.5415.2.2

publication LSID

lsid:zoobank.org:pub:081A068E-0352-4EB8-B45F-DFF23D9C811F

DOI

https://doi.org/10.5281/zenodo.10709195

persistent identifier

https://treatment.plazi.org/id/03DE87C4-211D-582D-A882-FAFAFA590D9F

treatment provided by

Plazi

scientific name

Anisoscelini
status

 

Key to the genera of Anisoscelini View in CoL (incl. Leptoscelini)

1. Hind tibia expanded, even if only weakly dilated with a flattened ridge limited to the outer surface [“ Anisoscelini View in CoL group”] ( Figs. 13, 15–18 View FIGURES 13–18 )........................................................................................... 2

1. Hind tibia simple or curved, perhaps quadrate in cross-section, but never dilated with a flattened lanceolate or foliate ridge [“Leptoscelini group”] ( Figs. 14 View FIGURES 13–18 , 26–31 View FIGURES 26–31 ).................................................................. 12

2. Hind tibia only weakly dilated with a flattened ridge limited to the outer surface ( Fig. 46 View FIGURES 44–49 )............................ 3

2’. Hind tibia conspicuously dilated, lanceolate or phylliform, at each side of the leg................................... 5

3. Corium hyaline, except the veins ( Figs. 16–17 View FIGURES 13–18 ); posterior margin of pronotal disk with a rounded triangular projection on either side of scutellar base ( Fig. 47 View FIGURES 44–49 )................................................. Holhymenia Lepeletier & Serville View in CoL

3’. Corium leathery, never hyaline; posterior margin of pronotal disk with..or without tiny triangular projections at either side of scutellar base........................................................................................ 4

4. Antennal segment I robust, shorter than or subequal to head, non-granulate; scutellum flat; femora ventrally armed with one or two acute subapical spines ( Fig. 30 View FIGURES 26–31 ).............................................................. Tarpeius Stål View in CoL

4’. Antennal segment I elongate, about 1.5x longer than head, cylindrical, irregularly granulate; scutellum with a depression near the apex; femora ventrally without acute subapical spine, usually with robust setae resembling a spine ( Fig. 13 View FIGURES 13–18 ).. Baldus Stål View in CoL

5. Anterior angles of pronotal disk projecting forward, lateral margins dentate; tylus spine-like ( Figs. 32 View FIGURES 32–37 , 48 View FIGURES 44–49 ).............................................................................................. Bellamynacoris Brailovsky View in CoL

5’. Anterior angles of pronotum without such projections; tylus usually obtuse (except Leptoglossus clypealis Heidemann View in CoL ).... 6

6. Body color dull-light-brown to dark-brown or black; never metallic, often less ostentatious species; light color limited to pale spots or stripe on the anterior pronotal disk, and/or a pale mid-corial fascia (zig-zag or straight), sometimes absent; antennal segments without flattened expansions.....................................................................7

6’. Body coloration highly variable, often with at least head or legs reddish to light-yellowish-orange (perhaps as stripes); usually metallic, colorfully ostentatious species; with or without pronotal marking and lacking a mid-corial fascia; antennal segments with or without flattened expansions...................................................................... 8

7. Antennal segment I shorter than preocular length, measurement from anterior border of eye to apex of tylus; rostral segment IV the longest ( Fig. 33 View FIGURES 32–37 )........................................................................... Narnia Stål View in CoL

7’. Antennal segment I longer than preocular length, measurement from anterior border of eye to apex of tylus; rostral segment I the longest ( Fig. 18 View FIGURES 13–18 )........................................................... Leptoglossus Guérin-Méneville View in CoL

8. Area between calli with two large tubercles; outer expansion of hind tibia lanceolate ( Fig. 34 View FIGURES 32–37 )........ Onoremia Brailovsky View in CoL

8’. Area between calli smooth, without tubercle; outer expansion of hind tibia phyliform................................9

9. Outer expansion of hind tibia as wide as the internal expansion; head, pronotum and scutellum metallic green with two yellowish longitudinal vittae extending from the apical jugum to nearly apex of the scutellum ( Fig. 15 View FIGURES 13–18 )............... Diactor Perty View in CoL

9’. Outer expansion of hind tibia always wider than inner expansion; lacking the combination of color above.............. 10

10. Outer expansion of hind tibia extending over most of tibial length and tapering apically; surface partially with green or bluish metallic iridescence ( Figs. 1–6 View FIGURES 1–6 )........................................................... Anisoscelis Latreille View in CoL

10’. Outer expansion of hind tibia short, not reaching the apex of tibia; surface without metallic iridescence (except Chondrocera mirabilis Brailovsky & Barrera View in CoL )......................................................................... 11

11. Antennal segment I twice as long as the head; hind femur with one row of acute spines ( Figs. 7-12 View FIGURES 7–12 )….......... Bitta Osuna View in CoL

11’. Antennal segment I less than twice as long as the head; hind femur with only two subapical spines ( Fig. 35 View FIGURES 32–37 )................................................................................................. Chondrocera Laporte View in CoL

12. Anterior pronotal calli with two conspicuous spines-like projections, each lateral to midline ( Fig. 49 View FIGURES 44–49 ); metallic-blue body with a bright-orange mid-corial fascia, alternating orange and blue connexival markings, and a mostly orange central pronotal disc ( Fig. 36 View FIGURES 32–37 ); known only from Cuba.................................................... Rhytidophthia Brailovsky View in CoL

12’. Anterior pronotal calli without two conspicuous spines-like projections; body coloration variable; broad geographic range... .................................................................................................. 13

13. Anterior angles of pronotal disk with triangular projections or spines directed anteriorly (or anterolaterally if part of a series of dentate lateral margin) ( Figs. 50–52 View FIGURES 50–56 )......................................................................14

13’. Anterior angles of pronotal disk rounded or straight, not projecting anteriorly or spinose, at most with a small tubercle ( Dalmatomammurius Brailovsky View in CoL )....................................................................... 18

14. Abdomen conspicuously expanded; anterolateral margins of pronotal disk moderately emarginated....................15

14’. Abdomen elongate, not expanded, or only weakly rounded laterally; anterolateral margins of pronotal disk not emarginated.....................................................................................................16

15. Tylus acutely projecting beyond and above the juga ( Fig. 50 View FIGURES 50–56 ); postocular tubercle well developed; antennal segment I elongate, robust, longer, length 1.90–2.21 mm; distance between antenniferous tubercle and anterior border of eye shorter (0.55–0.58 mm) ( Fig. 31 View FIGURES 26–31 )................................................................................ Ugnius Stål View in CoL

15’. Tylus not acutely projecting beyond and above the juga; postocular tubercle not visible; antennal segment I broad, shorter, length 1.04–1.09 mm; distance between antenniferous tubercle and anterior border of eye longer (0.79–0.81 mm) ( Fig. 37 View FIGURES 32–37 )........................................................................................... Plunentis Stål View in CoL

16. Mandibular plate well developed, close to the antenniferous tubercle; pronotal collar laterally with a short, acute, tooth-like tubercle on each side (when viewed dorsally); body elongate ( Fig. 38 View FIGURES 38–43 )....................... Kalinckascelis Brailovsky View in CoL

16’. Mandibular plate flat, not raised; pronotal collar laterally without a short acute spine on each side; body robust.......... 17

17. Pronotal disk piceous with reddish lateral margins, densely pubescent; antennal segments II and IV subequal; buccula slightly raised, evanescent; humeral angles obtuse, truncate, not laterally protruding ( Fig. 51 View FIGURES 50–56 ); posterolateral margins (behind humeri) of pronotal disk sinuate, not serrate or dentate ( Fig. 27 View FIGURES 26–31 ).................................... Phthiacnemia Brailovsky View in CoL

17’. Pronotal disk brown, largely glabrous; antennal segment II longer than IV; buccula quadrate, widely raised; humeral angles laterally protruding, apically with large to medium pointed projection ( Fig. 39 View FIGURES 38–43 ); posterolateral margins of pronotal disk serrate, even if weakly so ( Fig. 52 View FIGURES 50–56 )................................................................... Mamurius Stål View in CoL

18. Humeral angles of pronotal disk expanded or produced into a long acute spine or conspicuously foliate ( Fig. 53–54 View FIGURES 50–56 )...... 19

18’. Humeral angles of pronotal disk usually obtuse, rounded, and sometimes with a small subacute expansion ( Leptoscelis Laporte View in CoL ) ( Figs. 57, 59, 61 View FIGURES 57–62 )..................................................................................... 23 19. Body robust, larger species (27.80–32.00 mm)............................................................. 20

19’. Body elongate or slightly broad, smaller species (10.50–20.80 mm)............................................ 21

20. Humeral angles well-developed, wing expanded, dorsoventrally flattened and curving upward, coming to an acute spine with serrations anteriorly and posteriorly; anterior pronotal calli with two black tubercles at each side of midline; pronotum with a moderately produced but distinct transverse carina, between and behind the humeral angles; femora ventrally armed; yellowish-orange to dull-brown body, ventrally paler with black discoidal spot at mesopleuron, metapleuron and below each abdominal spiracle ( Fig. 22–25 View FIGURES 19–25 , 53–54 View FIGURES 50–56 )................................................................... Malvana Stål View in CoL

20’. Humeral angles not broadly expanded, but with a laterally projecting spine; pronotal calli without two black tubercles; anterior angles of pronotal disk densely granulate; femora unarmed; shiny orange and black body ( Fig. 40 View FIGURES 38–43 )... Sephinioides Brailovsky View in CoL

21. Humeral angles moderate to foliate, not produced into an acute spine; body always with green, bluish or purple metallic reflections; antennal segment IV conspicuously much longer than II (maximum length of II 5.00 mm, maximum length of IV 7.32 mm) ( Fig. 41 View FIGURES 38–43 ).................................................................. Malvanoides Brailovsky

21’. Humeral angles produced into a spine; body never with metallic reflections; antennal segment IV not much longer than II... .................................................................................................. 22

22. Pronotal humeral angles with acute, anteriorly projecting spines; body more elongate, parallel-sided; anterolateral margins of pronotal disk almost smooth; head clearly longer than wide; outer margin of antenniferous tubercle smooth ( Fig. 42 View FIGURES 38–43 )........................................................................................ Leptopelios Brailovsky View in CoL

22’. Pronotal humeral angles coming to a broad but sharp point; body robust; anterolateral margins of pronotal disk serrate; head as wide or wider than long; outer margin of antenniferous tubercles serrate ( Fig. 43 View FIGURES 38–43 )..................... Coribergia Casini View in CoL

23. Body always with green, bluish or purple metallic reflections................................................. 24

23’. Body without green, bluish or purple reflections............................................................ 26

24. Metathoracic peritreme undivided, not bilobate ( Fig. 58 View FIGURES 57–62 ); clavus and corium entirely with green metallic reflections ( Figs. 28 View FIGURES 26–31 , 57 View FIGURES 57–62 )................................................................................ Phthiadema Brailovsky View in CoL

24’. Metathoracic peritreme bilobate ( Figs. 60, 62 View FIGURES 57–62 ); clavus and corium rather dull or scattered with metallic reflections or entirely with greenish-blue metallic reflections................................................................... 25

25. Clavus and corium rather dull, except for a yellow transverse fascia; dorsal surface of head dull; anterolateral borders of pronotum smooth; abdominal sterna III–VI lateral to midline with rectangular yellowish to reddish-orange maculae ( Figs. 26 View FIGURES 26–31 , 55 View FIGURES 50–56 , 59 View FIGURES 57–62 )...................................................................................... Phthia Stål View in CoL

25’. Clavus and corium entirely with green to bluish reflections, unless with a yellow-orange transverse fascia sometimes expanded to occupy most of the clavus and corium; dorsal surface of head with green to bluish reflections; anterolateral borders of pronotum dentate; abdominal sterna III–VI lateral to midline without yellowish to reddish-orange maculae ( Figs. 29 View FIGURES 26–31 , 56 View FIGURES 50–56 , 61 View FIGURES 57–62 )................................................................................... Phthiarella Brailovsky View in CoL

26. Antennal segment I robust, shorter than total length of head................................................... 27

26’. Antennal segment I elongate, slender, longer than total length of head........................................... 28

27. Antenniferous tubercle with a blunt external spine ( Fig. 63 View FIGURES 63–65 ); anterior angles of pronotal disk acute, sometimes forming a tubercle; pronotal disk finely punctate, punctures not deeply excavated; tylus clearly raised above juga ( Fig. 14 View FIGURES 13–18 )..................................................................................... Dalmatomammurius Brailovsky View in CoL

27’. Antenniferous tubercle unarmed ( Fig. 64 View FIGURES 63–65 ); anterior angles of pronotal disk rounded; pronotal disk heavily punctate, punctures deeply excavated; tylus not raised above juga ( Fig. 44 View FIGURES 44–49 )........................................... Amblyomia Stål View in CoL

28. Body elongate, slender; dorsal body coloration dull-brown; antennal segment IV pale yellowish-orange; posterior half of hind femur armed dorsally with two rows of spines ( Fig. 45 View FIGURES 44–49 ).................................... Leptostellana Brailovsky View in CoL

28’. Body not slender, relatively broad; dorsal body coloration brown to black, often with contrasting markings of bright colors on the corium; antennal segment IV mostly red to yellow, if darker, then all veins of corium apically orange; posterior half of hind femur unarmed dorsally ( Figs. 19–21 View FIGURES 19–25 )..................................................... Leptoscelis Laporte View in CoL

Kingdom

Animalia

Phylum

Arthropoda

Class

Insecta

Order

Hemiptera

Family

Coreidae

SubFamily

Coreinae

Tribe

Anisoscelini

Genus

Malvana

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