Nugonioneuridae Carpenter, 1976

Aristov, D. S., 2020, Revision of the genera Heteroptilon, Nugonioneura and Opistocladus from the Lower Permian of USA (Insecta: Cnemidolestida: Tillyardembiidae, Nugonioneuridae and Parmapteridae), Far Eastern Entomologist 401, pp. 1-9 : 5

Family Nugonioneuridae Carpenter, 1976 , nom. resurr.

Nugonioneuridae Carpenter, 1976: 377 (type genus: Nugonioneura Tillyard, 1937 ); Rasnitsyn, 1980: 43; Carpenter, 1992: 500; synonimized under Tococladidae by Rasnitsyn

(2002).

Type genus: Nugonioneura Tillyard, 1937 .

DIAGNOSIS. Small insects. Forewings preradial field is moderately wide. SC

ends in forking into C and R near wings mid length. RS starts at the wing’s mid length, drawn near to SC apex, merged with MA, equal in size with CuA. Base of M

at the short area merged with CuA, M starts branching at wings mid length. CuA without posterior branches, starts branching before its mid length. Apices of anal veins are not merging with each other.

COMPARISON. Nugonioneura is most similar to Psoroptera Carpenter, 1976

( Cnemidolestida : Psoropteridae ) (Aristov, 2014) from the Permian locality Elmo in

USA, it differs in SC, ends with forking into C and R, apex of SC drawn close to base of RS, which starts at wings mid length, lacking of anal loop. In Psoropteridae

SC ends at C, apex of SC not drawn close with base of RS, which starts at wings basal quarter (Aristov, 2014).

GENERAE INCLUDED. Type genus.

NOTE. Like Heteroptilon , Nugonioneura was attributed to hypoperlids Tococladidae (Rasnitsyn, 2002) . Heteroptilon differs from the most similar genus

Opistocladus in late branching of M, anastomosis of M and CuA bases and lacking of posterior branches of CuA. In Tococladidae M branching in wings basal third, M

and CuA are connected by М 5, CuA have posterior branches in intercubital field

(Carpenter, 1976; Béthoux et al., 2003).