Acylophorus madecassus, Janák, 2018

Janák, Jiří, 2018, Revision of the genus Acylophorus Nordmann, 1837 from Madagascar and Mascarene Islands (Coleoptera: Staphylinidae: Staphylininae: Acylophorina), Zootaxa 4462 (2), pp. 151-191 : 167

Acylophorus madecassus sp. nov.

( Figs. 67–77 , 175 )

Type locality. Madagascar, Sambava , near Anovona river, cca 14 15' 54" S 50 09' 02" E.

Type material. Holotype ♂: MADAGASCAR: Madagascar Est , Sambava , Riv. [Rivière = river] Anovona env., 0–20 m, 3.– 4.3.1996, rizières/bord de la forêt [rice fields/forest edge], J. Janák & P. Moravec lgt. ( JJRC) .

Description. Body length 7.5 mm, forebody length 3.4 mm. Body black ( Fig. 67 ), pronotum and abdomen brown, abdominal tergites markedly iridescent; antennae brown with base of first two segments and segments 7–11 reddish yellow; legs reddish brown, tarsi and maxillar palpi reddish yellow, abdomen gold iridescent.

Head subrounded ( Fig. 68 ), about as long as wide (L/W=1.01), markedly widened behind eyes, with two pairs of interocular setae. Temples longer than eyes (R=1,56). Dorsal side of head shining, without scatered extremely fine micropunctures. Dense pale grey pubescece behind eyes. Four postocular setae visible on each side, additional seta on hind margin of eye present. Left mandible with two teeth, right mandible with three teeth ( Fig. 69 ). Maxillary palpi with terminal segment long and relativelly narrow, pubescent, asymmetric, markedly wider and longer than glabrous penultimate segment which is elongate ( Fig. 70 ). First segment of antenna nearly as long as next four; segments 1 to 4 elongate, segment 5 markedly longer than wide (R=1.61), segment 8 about as long as wide, segment 9 markedly transverse, segment 10 markedly transverse (R=0.62).

Pronotum slightly transverse (R=1.14), markedly widened behind with sides rounded, widest in basal half ( Fig. 68 ), shining, with scattered extremely fine micropunctures. One pair of dorsal setae and one pair of lateral setae. Marginal setae short. Elytra slightly transverse (R=1.36) with short, pale, but not shining pubescence arising from dense aspirate punctures. Apical fringe of bristles markedly longer than pubescence on other parts of the elytra. Abdominal tergites with moderately long and moderately sparse pubescense, arising from aspirate punctures that are irregularly distributed across each tergite and more coarse but markedly sparser than on elytra.

Meso- and metatarsi with empodial setae longer than claws (similar as in Figs. 9, 10 ).

Male. Sternite 9 very long and narrow ( Fig. 77 ), apex of tergite 10 narrowly rounded, with very long setae ( Fig. 76 ). Aedeagus ( Figs. 71–75 ) 0.92 mm long. Parameres bilobed, lobes widely separated, narrowed apically, sensory pegs confusedly arranged in apical half of both lobes ( Fig. 75 ). Median lobe of aedeagus markedly longer than paramere, apex broadly rounded ( Figs. 73 , 75 ).

Female. Unknown.

Differential diagnosis. Acylophorus madegassus sp. nov. differs from all hirtherto known Aftrotropical Acylophorus by the very long temples and by the different shape of aedeagus.

Figures

FIGURES 67–73. Acylophorus madecassus sp. nov., holotype. 67, habitus; 68, head and pronotum; 69, mandibles; 70, maxillary palpus; 71, aedeagus lateral; 72, aedeagus ventro-lateral; 73, aedeagus ventral. 71–73, same scale bars.

FIGURES 74–77. Acylophorus madecassus sp. nov., holotype. 74, aedeagus lateral; 75, aedeagus ventral; 76, male tergites 9, 10; 77, male sternite 9. 74, 75, transmitted light, same scale bars.

FIGURES 172–177. Distribution maps. 172, Acylophorus tristis Jarrige; 173, A. derougemonti sp. nov.; 174, A. muscorum Jarrige; 175, A. delphinus Fauvel (D), A. madecassus sp. nov. (Ŧ); 176, A. minutus Jarrige (D), A. hova sp. nov. (Ɨ), A. montanus sp. nov. (Δ); 177, A. sogai Jarrige.

FIGURES 9–17. Acylophorus tristis Jarrige. 9, 10, 15–17, Col de Marotaolana; 11–13, Sendrisoa; 14, Manindray. 9, metatarsus lateral; 10, metatarsus dorsal; 11, 16, aedeagus lateral; 12, aedeagus ventro-lateral; 13–15, aedeagus ventral. ES, empodial setae. 16, 17, transmitted light. 9–10, 11–17, same scale bars.