Tenuosmylus brevineurus, Wang & Liu & Ren, 2009
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publication ID |
https://doi.org/ 10.4202/app.2008.0040 |
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persistent identifier |
https://treatment.plazi.org/id/03DCBB53-5A05-FFDC-FCC9-FB65FDEB38B6 |
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treatment provided by |
Felipe |
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scientific name |
Tenuosmylus brevineurus |
| status |
sp. nov. |
Tenuosmylus brevineurus View in CoL sp. nov.
Figs. 2 View Fig 2 , 3 View Fig 3 .
Etymology: The name is a combination of the Latin brevis (meaning, “short”) and neurus (meaning, “vein”) referring to the short Cu 2 in the forewing.
Holotype: CNU−NN99030. Specimen consists of a relatively complete body with four wings spread out. Only the left forewing provides discernable venation.
Type locality: Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China.
Type horizon: Jiulongshan Formation, Aalenian–Bajocian, Middle Jurassic.
Material.— Holotype CNU−NN99030 and paratype CNU−NN99031 (specimen with head and thorax anamorphic. Four wings folded posteriorly).
Diagnosis.—The same as for the genus (monotypic at present).
Description.—Eyes distinctly dark. Antennae filiform but length uncertain due to poor preservation, scape slightly bulky. Prothorax trapezoid. Meso− and metathorax dilated. Mesonotum equipped with complete sclerites, prescutum small and semicircular, scutellum triangular. Metanotum with fully−developed sclerites. Abdomen poorly preserved with only an inconspicuous trace.
Sc R1
A1
Forewing elongated with subacute apex, 18.1 mm long, 4.5 mm wide. Trichosors present but inapparent at costal margin, perhaps due to poor preservation. Pterostigma distinctly fuscous. Membrane colorless, and nygmata undetected. Costal space narrow at base and then dilated. Subcostal space very narrow. Sc and R1 fused distally and entering the margin before the apex, sc−r absent. Rs originating close to wing base. Numerous cross−veins present between R1 and Rs. Rs with seven branches, each forked distally. Mp forked near wing base with two parallel branches that are each distally bifurcated. A1 not well preserved but also with pectinate branches. A2 and A3 inapparent.
Hindwing poorly preserved with trichosors inapparent. Costal space narrow with simple veinlets lacking distal forks. Subcostal space extremely narrow, sc−r not visible. Cross−veins between R1 and Rs numerous. Separation of Ma and first branch of Rs similar to in forewing. Mp forked close to wing base, the distance between the two branches slightly dilated in middle. Mp2 bifurcated distally. Anal veins not well preserved, only a single fragmentary one is apparent.
Discussion.—Our newly described genus is being assigned to Gumillinae based on defining characteristics of the subfamily. This includes the following combination of characters: excessively long antennae (poorly preserved in the specimens); greatly elongated wings; trichosors limited at outer and posterior margins and partly at costal margin; narrow costal space with simple costal cross−veins; separation of Ma from Rs generally distant from wing base (at nearly one−third of wing length); first branch of Rs near middle of wing; presence of more than three presectoral cross−veins; branches of Mp distally forking as they enter outer wing margin; distal branches of Cu1 pectinate or deeply forked, branches of Cu2 pectinate from middle to terminal end; A1 parallel with Cu in forewing; hind wing with a single Cu branch. We also concur with Menon and Makarkin (2008) in the placement of four other genera ( Nymphites Hasse, 1890 ; Epiosmylus Panfilov, 1980 ; Enodinympha Ren and Engel, 2007 ; Nilionympha Ren and Engel, 2007 ) into this subfamily.
Stratigraphic and geographic range.—Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China. Jiulongshan Formation, Middle Jurassic.
Acknowledgements.—We sincerely thank Shih ChungKun (College of Life Science, Capital Normal University, Beijing, China) for revision of the manuscript. We are grateful to Roy Vogtsberger (Department of Biology, Hardin−Simmons University, Abilene, USA), Andre Nel (Muséum National d’Histoire Naturelle, Paris, France) and Rafael Gioia Martins Neto (Universidade Federal de Juiz de Fora, Juiz de Fora, Brazil) for their improvement of our manuscript. The research has been supported by grants from the National Nature Science Foundation of China (No. 30430100, 40872022), the Nature Science Foundation of Beijing (No.5082002) and Scientific Research Key Program and PHR Project of Beijing Municipal Commission of Education.
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