HARPALINAE, Bonelli, 1810

Peck, Stewart B., Thomas, Michael C. & Robert H. Turnbow, Jr., 2014, The diversity and distributions of the beetles (Insecta: Coleoptera) of the Guadeloupe Archipelago (Grande-Terre, Basse-Terre, La Désirade, Marie-Galante, Les Saintes, and Petite-Terre), Lesser Antilles, Insecta Mundi 2014 (352), pp. 1-156 : 15-18

SUBFAMILY HARPALINAE

TRIBE HARPALINI

SUBTRIBE HARPALINA

SELENOPHORINI GROUP

Amblygnathus cephalotes Dejean 1829: 63 ; Erwin and Sims 1984: 441; Ball and Maddison 1987: 245; Ball 1992: 82. = Amblygnathus vitraci Fleutiaux and Sallé 1890: 364 of Guadeloupe. Distribution. Dominica, Guadeloupe. Bolivia, French Guiana, Surinam; Lesser Antilles and Latin America .

Amblygnathus gilvipes Ball and Maddison 1987: 230 , 232. Distribution. Guadeloupe. Surinam to Brazil and Bolivia; Lesser Antilles and Latin America .

Athrostictus iridescens Chaudoir 1843: 783 ; Erwin and Sims 1984: 441. Distribution. Guadeloupe; single island endemic.

Discoderus subaeneus Reiche 1843: 141 ; Fleutiaux and Sallé 1890: 366 ( Selenophorus ); Erwin and Sims 1984: 441. Distribution. Guadeloupe. Panama, South America ; Lesser Antilles and Latin America .

Gynandropus guadeloupensis Fleutiaux and Sallé 1890: 365 ; Wolcott 1950: 229 ; Erwin and Sims 1984: 440 (as Selenophorus ); Ball and Maddison 1987: 232. Distribution. Guadeloupe, Puerto Rico; widespread Antilles endemic.

Selenophorus alternans Dejean 1829: 49 ; Fleutiaux and Sallé 1890: 366; Ramos 1946: 31; Ivie et al. 2008b: 238; Valentine and Ivie 2005: 275; Turnbow and Thomas 2008: 14. = Selenophorus macleayi (Kirby) 1837: 50 ; Blackwelder 1944-1957: 49; Bennett and Alam 1985: 20; Ball 1992: 85. Distribution. Bahamas, Barbados, Cayman Islands, Cuba, Guadeloupe, Guana, Hispaniola, Jamaica, Mona, Montserrat, Puerto Rico. Mexico to Venezuela to Uruguay; widespread Antilles and Latin America .

Selenophorus chalybaeus Dejean 1829: 110 ; Fleutiaux and Sallé 1890: 366; Ball 1992: 85; Ivie et al. 2008b: 238; Turnbow and Thomas 2008: 14; Perez-Gelabert 2008: 79. Distribution. Antigua, Bahamas, Dominica, Guadeloupe, Hispaniola, Montserrat, Puerto Rico. Central and South America ; widespread Antilles and Latin America .

Selenophorus propinquus Putzeys 1874: 118 ; Fleutiaux and Sallé 1890: 366; Erwin and Sims 1984: 440; Ball 1992: 85; Valentine and Ivie 2005: 275; Ivie et al. 2008b: 238; Turnbow and Thomas 2008: 14. Distribution. Antigua, Bahamas, Dominica, Guadeloupe; Guana, Montserrat. South America ; Lesser Antilles and Latin America . Notes. Records of Selenophorus integer (Fabricius) 1801: 196 ( Harpalus ) in Fleutiaux and Sallé 1890: 366 and Erwin and Sims 1984: 440 ( Harpalus )) for Lesser Antilles are this species; Selenophorus integer is confined to the Greater Antilles.

Selenophorus sinuatus Gyllenhal 1806: 203 ; Fleutiaux and Sallé 1890: 366; Ramos 1946: 31; Erwin and Sims 1984: 441; Ball 1992: 85; Valentine and Ivie 2005: 275; Ivie et al. 2008b: 238; Turnbow and Thomas 2008: 15. Distribution. Antigua, Bahamas, Cayman Islands, Cuba, Dominica, Guadeloupe, Guana, Hispaniola, Jamaica, Les Saintes, Mona, Montserrat, Puerto Rico; widespread Antilles endemic.

Selenophorus subquadratus Putzeys 1878: 59 ; Erwin and Sims 1984: 441; Ball 1992: 85; Ivie et al. 2008b: 238. = Selenophorus guadeloupensis Fleutiaux and Sallé 1889: 365 ; Ball 1992: 96. Distribution. Cuba, Dominica, Guadeloupe, Hispaniola, Montserrat, unspecified Windward Islands; widespread Antilles endemic.

TRIBE PENTAGONICINI

Pentagonica flavipes flavipes (LeConte) 1853: 377 ( Didetus ); Reichardt 1968; Bell 1985: 323; Miskimen and Bond 1970: 79; Schiller 2004: 34; Valentine and Ivie 2005: 275; Ivie et al. 2008b: 238; Turnbow and Thomas 2008: 14; Perez-Gelabert 2008: 81.= Rhombodera picea Fleutiaux and Sallé 1890: 362 of Guadeloupe. Distribution. Antigua, Bahamas, Cuba, Dominica, Grand Cayman, Guadeloupe, Guana, Hispaniola, Montserrat, St. Croix. SE USA, Mexico , Central America , Colombia, Trinidad, Brazil; widespread New World. The subspecies P. f. picipes Darlington occurs on Hispaniola, Jamaica, Puerto Rico, and St. Croix.

TRIBE PLATYNINI

Dyscolus spp. - ellipticus species group; restricted to Lesser Antilles, flight wings completely or partially reduced

Dyscolus alternans (Chaudoir) 1878: 348 ( Colpodes ); Fleutiaux and Sallé 1890: 368; Liebherr, 1988: 138 ( Platynus ). Distribution. Guadeloupe; single island endemic.

Dyscolus lherminieri (Chaudoir) 1842: 838 ( Paranomus ), Chaudoir 1859: 318 ( Colpodes ); Fleutiaux and Sallé 1890: 367; Liebherr 1987: 352 ( Platynus ), 1988: 138; Schiller 2004: 43. Distribution. Dominica, Guadeloupe; Lesser Antilles endemic.

Dyscolus spp. - memnonius species group; also occurring in Greater Antilles and on continental mainland (winged)

Dyscolus dejeani (Chaudoir) 1859: 359 ( Colpodes ( Dyscolus )); Fleutiaux and Sallé 1890: 367; Liebherr, 1987: 355, 1988: 138 ( Platynus ). = Platynus brunnea Dejean 1831: 440 of Guadeloupe. Distribution. Guadeloupe; single island endemic.

Dyscolus elongatus (Chaudoir) 1878: 344 ( Colpodes ); Fleutiaux and Sallé 1890: 367; Liebherr 1987: 356, 1988: 138 ( Platynus ). Distribution. Guadeloupe; single island endemic.

Dyscolus memnonius Dejean 1831: 439 ; Chaudoir 1878: 343 ( Colpodes ); Fleutiaux and Sallé 1890: 367; Liebherr 1987: 358, 1988: 138 ( Platynus ). Distribution. Guadeloupe; single island endemic.

Glyptolenus chalybaeus Dejean 1831: 720 ; Fleutiaux and Sallé 1890: 368 ( Colpodes ); Erwin and Sims 1984: 435 ( Platynus ); Liebherr 1997: 90; Ivie et al. 2008b: 238. Distribution. Dominica, Grenada, Guadeloupe, Montserrat, St. Cristopher, St. Vincent. Costa Rica, Nicaragua, Panama, Brazil; Lesser Antilles and Latin America .

TRIBE PERIGONINI

Perigona guadeloupensis Fleutiaux and Sallé 1890: 367 ; Erwin and Sims 1984: 443. Distribution. Guadeloupe; single island endemic.

Perigona nigriceps Dejean 1831: 44 ; Erwin and Sims 1984: 443; Bennett and Alam 1985: 20; Valentine and Ivie 2005: 275; Ivie et al. 2008b: 238; Turnbow and Thomas 2008: 14; Perez-Gelabert 2008: 81. Distribution. Bahamas, Barbados, Cuba, Dominica, Guadeloupe, Guana, Hispaniola, Martinique, Montserrat, Puerto Rico. Canada (PQ), USA (NH-FL-CA); introduced to Lesser Antilles. Note. Introduced to New World; an Old World (probably Asian) species widely distributed by commerce in the New World. Found around human dwellings, and probably living in decaying plant matter in and around gardens.

Perigona picea Darlington 1934: 98 ; Erwin and Sims 1984: 443. Distribution. Cuba, Hispaniola, Guadeloupe; widespread Antilles endemic.

TRIBE LACHNOPHORINI

Anchonoderus subaeneus Reiche 1843: 40 ; Liebherr 1988: 27. Distribution. Guadeloupe, probably introduced. Ecuador, Colombia, French Guiana, Guatemala, Panama; introduced to Lesser Antilles?

TRIBE LEBIINI

SUBTRIBE APENINA

Apenes spp. - purpurata species group; known only from the Greater and Lesser Antilles, except Puerto Rico.

Apenes chalumeaui Ball and Shpeley in Ball 1992: 115; Ivie et al. 2008b: 238; Ball and Shpeley 2009: 130. Distribution. Guadeloupe (type locality), Montserrat, St. Kitts; Lesser Antilles endemic.

Apenes purpurata Fleutiaux and Sallé 1890: 36 ; Erwin and Sims 1984: 445; Ball 1992: 88; Ball and Shpeley 2009: 128. Distribution. Dominica, Guadeloupe (type locality); Lesser Antilles endemic.

Apenes spp. - sculpticeps species group; a wide ranging group from Brazil to Mexico , with only one species known in the West Indies.

Apenes sculpticeps Ball and Shpeley 2009: 121 = Apenes marmorata Ball 1992: 106 , not Chaudoir 1875: 41. Distribution. Guadeloupe, St. Croix. Honduras to Brazil; widespread Antilles and Latin America .

Apenes spp. - variegata species group; wide ranging from sw USA to Argentina and on most West Indian islands

Apenes marginalis (Dejean) 1831: 315 ( Cymindis ); Fleutiaux and Sallé 1890: 360; Miskimen and Bond 1970: 78; Erwin and Sims 1984: 445; Bennett and Alam 1985: 20; Ivie et al. 2008b: 238; Ball and Shpeley 2009: 106. Distribution. Anguilla, Barbados, Dominica, Grenada, Guadeloupe, Hispaniola, Jamaica, Les Saintes, Martinique, Montserrat, Mustique, Puerto Rico, St. Croix, St. Lucia, St. Vincent, Turks and Caicos (Grand Turk). Southwestern USA to Honduras, Panama, Trinidad, Brazil; Lesser Antilles and Latin America . Notes. Probably a predator on pests attacking crop plants.

Apenes variegata (Dejean) 1825: 217 ( Cymindis ); Erwin and Sims 1984: 44, as synonym of Apenes pallipes (Fabricius) ; Ball 1992: 88; Ball and Shpeley 2009: 104. = Cymindis guadeloupensis Gory 1833: 196 . Distribution. Anguilla, Antigua, Desirade, Guadeloupe, Les Saintes, Marie-Galante, Mustique, Puerto Rico, St. Barthélemy (type locality), St. Eustatius, St. John, St. Kitts, St. Martin, St. Vincent. Trinidad, Brazil, to n Mexico ; widespread Antilles and Latin America . Note. Lesser Antilles records (e. g. Fleutiaux and Sallé 1890: 360 as Cymindis ) for Apenes pallipes (Fabricius) 1792: 159 ( Carabus ), which is a North American species of Agonum , are of this species.

SUBTRIBE LEBIINA

Lebia frenata Chaudoir 1871: 171 ; Erwin and Sims: 1984: 447. = Aphelogenia frenata apicalis Fleutiaux and Sallé 1890: 36 l of Guadeloupe. Distribution. Guadeloupe (subspecies Lebia frenata chevrolati Blackwelder 1944 -57: 54, replacement name for junior homonym Lebia apicalis ). French Guiana; Lesser Antilles and Latin America .

SUBTRIBE PERICALINA

Somotrichus unifasciatus Dejean 1831: 7 ; Erwin and Sims 1984: 446. = Lebia elevata Fabricius 1787: 198 in Fleutiaux and Sallé 1890: 361. Distribution. Guadeloupe. USA; cosmopolitan; introduced to New World; introduced to Lesser Antilles.

TRIBE GALERITINI

Galerita americana (L.) 1758: 415 ( Carabus ); Fleutiaux and Sallé 1890: 359; Reichardt 1967: 63. = Galerita geniculata Dejean 1831: 297 of Guadeloupe. Distribution. Guadeloupe, St. Martin. Widespread from Guatemala to Panama, to Trinidad, south to Brazil, Paraguay, and Bolivia; widespread Antilles and Latin America .

Galerita tristis Reiche 1842: 273 ; Erwin and Sims 1984: 442; Reichard 1967: 96. = Galerita unicolor Dejean 1824: 117 in Fleutiaux and Sallé 1890: 359 of Guadeloupe. Distribution. Barbados, Dominica, Guadeloupe, Jamaica. Costa Rica, El Salvador, Panama, South America , Trinidad; widespread Antilles and Latin America .