Charipinae, Dalla Torre & Kieffer, 1910

Ferrer-Suay, Mar, Selfa, Jesús & Pujade-Villar, Juli, 2018, Palaearctic species of Charipinae (Hymenoptera, Figitidae): two new species, synthesis and identification key, European Journal of Taxonomy 427, pp. 1-110 : 4-12

Key of Palaearctic Charipinae species

1. Metasoma with a single tergal plate, or if two, then basal tergite much shorter than second along middorsal line (Fig. 1.4) ……………………………………………………………………………2

– Metasoma with two large visible terga, subequal in length along middorsal line, but basal tergite ¼–½ smaller than second in lateral view (Fig. 1.3) ……………………………………………7

2. Basal tergite much shorter than second along middorsal line ………… Apocharips Fergusson, 1986 Radial cell short, 1.2 times as long as wide, with R1 and Rs parallel ……………………………… ……………………………………………………………… Apocharips trapezoidea ( Hartig, 1841)

– Metasoma with a single tergal plate …………………………………………3 ( Dilyta Förster, 1869 )

3. Metasoma without punctures …………… Dilyta aleevae Pujade-Villar & Paretas-Martínez, 2011 – Metasoma with a punctate area on distal part ………………………………………………………4

4. Male: F1 very long, wide and arched, much longer than pedicel (almost double) and longer than F2 and F3 together;F2slightly shorter or subequal to F3;F4 longer than F2 and F3; F4–F12wider than previous segments; antenna slightly clavate from F4; sensilla beginning on F4 (Fig. 6.5). Female unknown … ………………………………………………… Dilyta sinica Ferrer-Suay & Paretas-Martínez, 2011

– Male, when known, with different features than given above ………………………………………5

5. Female: F1 very long, thin, almost twice as long as pedicel, longer than F2–F5 and nearly as long as F2+F3+F4 combined (Fig. 6.2). Male unknown ……………………………………………… ……………………………………………… Dilyta japonica Paretas-Martínez & Ferrer-Suay, 2011

– F1 subequal to pedicel and shorter or subequal to F2 and F3 combined ………………………………6

6. Female: F1 slightly shorter or subequal than pedicel; F2 subequal to F3; F4 slightly shorter than F1, but longer than F2 or F3; F1 subequal to F5; F6 longer than F5 (Fig. 6.6). Male: F1 slightly longer than pedicel; F2 or F3 each shorter than F1; F1 subequal to F4; F4–F12 wider than previous flagellomeres; antenna slightly clavate from F4; sensilla beginning on F4 ……… Dilyta subclavata Förster, 1869

– Female: F1 subequal to pedicel or slightly longer; F2 shorter than F3; F3 shorter than F4; F4 shorter than F5; F1 subequal to F5 (Fig. 6.3). Male: F1 subequal to pedicel; F2 shorter than F1 or F3; F3 subequal to F1; F3–F12 wider than previous flagellomeres; antenna slightly clavate from F3; sensilla beginning on F3 …… Dilyta longinqua Paretas-Martínez & Pujade-Villar, 2011

7. Lower part of mesopleuron with horizontal sulcus (Fig. 1.1) …8 ( Phaenoglyphis Förster, 1869 ) – Mesopleuron lacks horizontal sulcus (Fig. 1.2) ………………………23 ( Alloxysta Förster, 1869 )

8. Notauli present, at least in the posterior half of mesoscutum and/or scutum sculptured ……9 – Notauli completely absent and scutum smooth and shining ……………………………………21

9. Mesoscutum covered by imbricate sculpture (except mesopleura) ………………………………10 – Mesoscutum smooth, without imbricate sculpture ………………………………………………12

10. Mesoscutum mostly smooth, with a few wrinkles on the distal side of the notauli; rhinaria and club shape begin on the last three quarters of F1 (Fig. 6.18). Male antenna: rhinaria and club shape begin on F1; F1 curved; F1 longer than pedicel and F2; F2–F11 subequal ………………………… …………………………………………………………… Phaenoglyphis ruficornis ( Förster, 1869)

– With distinctive imbricate sculpture on all surfaces; rhinaria and club shape begin on different flagellomere. Male unknown ………………………………………………………………………11

11. Rhinaria and club shape begin on F1 (Fig. 6.17); thick pronotal carinae not reaching the mesoscutum; mesoscutum enterely covered by many setae; notauli present, only indicated anteriorly and well-marked posteriorly; scutellar foveae open posteriorly (Fig. 7.11); central part of metascutellum smooth, only with a central carina; fore wing with long marginal setae. Male unknown ……………………………………………… Phaenoglyphis pubicollis ( Thomson, 1877)

– Rhinaria and club shape begin on F4 (Fig. 6.8); long pronotal carinae reaching the mesoscutum; mesoscutum with few scattered setae; notauli only indicated; scutellar foveae completely defined and with a transverse posterior carina inside (Fig. 7.2); central part of metascutellum with imbricate sculpture; fore wing without marginal setae (Fig. 10.1). Male unknown ……………… ………………………………… Phaenoglyphis evenhuisi Pujade-Villar & Paretas-Martínez, 2006

12. Head, mesosoma and metasoma yellowish brown; notauli deeply excavate, rounded scutellar foveae, but with interior side straight (Fig. 7.16). Male unknown …………………………………… ………………………………………………………… Phaenoglyphis xanthochroa Förster, 1869

– Head, mesosoma and metasoma dark brown; notauli present but not deeply excatave; different shape of scutellar foveae …………………………………………………………………………………13

13. Antenna longer than body …………………………………………………………………………14 – Antenna subequal to or shorter than body ………………………………………………………18

14. Rhinaria and club shape begin on F1 ………………………………………………………………15 – Rhinaria and club shape begin on other flagellomeres ……………………………………………17

15. Rhinaria and club shape begin at the proximal end of F1; F2 subequal to F3 and F3 shorter than F4 (Fig. 6.13); Rs slightly curved. Male unknown … Phaenoglyphis longicornis ( Hartig, 1840)

– Rhinaria and club shape begin on the last three-quarters of F1; different proportions between flagellomeres; Rs straight …………………………………………………………………………16

16. Last flagellomere 2.5 times as long as wide (Fig. 6.20); scutellar foveae with straight sides and open anteriorly and posteriorly (Fig. 7.14); radial cell 2.4 times as long as wide. Male unknown … …………………………………………………………… Phaenoglyphis stricta ( Thomson, 1877)

– Last flagellomere 4.3 times as long as wide (Fig. 6.11); rounded scutellar foveae, slightly open posteriorly (Fig. 7.5); radial cell 2.9 times as long as wide. Male unknown ……………………… …………………………………………………………… Phaenoglyphis insperatus Belizin, 1973

17. Rhinaria and club shape not beginning on the same flagellomere, rhinaria begin on F1 and club shape begins on F3, F1 longer than pedicel (Fig. 6.9). Male antenna: club shape begins on F3, rhinaria on F5, F2 subequal to F3, F3 shorter than F4 ……………………………………………… ……………………………………………………… Phaenoglyphis fuscicornis ( Thomson, 1877)

– Rhinaria and club shape both begin on F3, F1 shorter than pedicel (Fig. 6.10). Male antenna: rhinaria and club shape begin on F3, F2 shorter than F3, F3 subequal to F4 ……………… …………………………………………………………… Phaenoglyphis heterocera ( Hartig, 1841)

18. F1 longer than pedicel, 1.1 times as long as pedicel and longer than F2; F2 shorter than F3; F3– F4 subequal in length (Fig. 6.19); mesoscutum with a line of setae next to each notaulus; notauli weakly present (Fig. 7.13); scutellar foveae completely defined and with two lines anteriorly; propodeum with two short and straight carinae reaching the base independently. Male unknown ……………………………………………………………… Phaenoglyphis salicis ( Cameron, 1883)

– F1 subequal to or shorter than pedicel; different combination of features …………………19

19. Beginning of rhinaria in F2; F1 longer than F2; F2–F4 subequal in length; radial cell 2.7 times as long as wide …………………………………………… Phaenoglyphis proximus Belizin, 1966

– Beginning of rhinaria in F3; different proportions between flagellomeres; different size of radial cell …………………………………………………………………………………………………20

20. Female: F1 subequal to F2; F2–F4 subequal in length (Fig. 6.14); scutellar foveae rounded and separated by a thin carina (Fig. 7.8); with abundant setae on apex of scutellum; propodeum with narrow carinae (sometimes difficult to see). Male antenna: rhinaria and club shape begin on F3; F1 not curved, longer than pedicel and F2; F2 shorter than F3; F3–F4 subequal ………… …………………………………………………………… Phaenoglyphis moldavica Ionescu, 1969

– Female: F1 longer than F2, but shorter than F3; F3 subequal to F4 (Fig. 6.7); scutellar foveae completely defined (Fig. 7.1). Male unknown …… Phaenoglyphis abbreviata ( Thomson, 1877)

21. Scutellar foveae not present …………………………… Phaenoglyphis nigripes ( Thomson, 1877) – Scutellar foveae present, sometimes superficially …………………………………………………22

22. Radial cell partially open along anterior margin; F1 and F2 subequal; F1 curved in males (Fig. 6.21) …………………………………………………… Phaenoglyphis villosa ( Hartig, 1841)

– Radial cell closed; F1 longer than F2 (Fig. 6.12). Male unknown …………………………… …………………………………………………………… Phaenoglyphis insularis ( Belizin, 1973)

23. Brachypterous species ……………………………………………………………………………24 – Fully winged species, wing usually longer than mesosoma and metasoma ………………………31

24. Fore wing reaching the end of the metasoma; visible radial cell …………………………………25 – Fore wing reaching the beginning of the metasoma; without radial cell visible ……………………28

25. Completely open radial cell (Fig. 4.22) …………………… Alloxysta marshalliana ( Kieffer, 1900) – Closed radial cell …………………………………………………………………………………26

26. Pronotal carinae absent …………………………………………… Alloxysta glebaria Hellén, 1963 – Pronotal carinae present ……………………………………………………………………………27

27. Propodeal carinae absent ………………………… Alloxysta pseudoconsobrina Ferrer-Suay, 2017 – Propodeal carinae present ……………………………………… Alloxysta curta Ferrer-Suay, 2017

28. Pronotal carinae present; propodeal carinae absent.Male always and female sometimes brachypterous ……………………………………………………………… Alloxysta halterata ( Thomson, 1862) – Pronotal carinae absent; propodeal carinae absent or present ………………………………………29

29. Propodeal carinae well defined, joining posteriorly. Male: F1 shorter than pedicel (Fig. 2.8). Female unknown …………………………………………… Alloxysta brachyptera ( Hartig, 1840) – Propodeal carinae absent. F1 shorter or longer than pedicel ……………………………………30

30. Fore wing reaching the beginning of metasoma. Female: F1 longer than pedicel and F2; F2 subequal to F3 (Fig. 3.1). Male: F1 subequal or slightly longer than pedicel, F1–F4 subequal in length ………………………………………………………… Alloxysta pedestris ( Curtis, 1838)

– Fore wing very short, practically absent. Female: F1 shorter than pedicel, F1 longer than F2, F2 subequal to F3 (Fig. 2.3). Male unknown ……… Alloxysta apteroidea Hellén, 1963

31. Radial cell open or partially open ………………………………………………………………32 – Radial cell closed …………………………………………………………………………………58

32. Radial cell completely open ………………………………………………………………………33 – Radial cell partially open …………………………………………………………………………43

33. Propodeal carinae absent …………………………………………………………………………34 – Propodeal carinae present …………………………………………………………………………40

34. Male: F1 and F2 clearly bumped; F1 subequal to F2; F2 longer than F3, but shorter than F4 (Fig. 11.2–3); radial cell 2.3 times as long as wide (Fig. 11.4). Female unknown …… ……………………………………… Alloxysta palearctica Ferrer-Suay & Pujade-Villar sp. nov.

– Male (when known) with F1 and F2 not bumped ………………………………………………35

35. F2 longer than F1 and F3 (Fig. 3.6); radial cell 2.3 times as long as wide (Fig. 5.10) ……… …………………………………………………………………… Alloxysta proxima Belizin, 1962 – F2 shorter than or subequal to F1; radial cell longer or shorter than 2.3 times as long as wide …36

36. Female: F2 subequal to F1; F2 longer than F3 (Fig. 2.6); radial cell 3.0 times as long as wide (Fig. 4.5). Male unknown ………………………………… Alloxysta basimacula ( Cameron, 1886)

– Female: F2 shorter than F1, F2 longer, shorter than or subequal to F3; radial cell longer or shorter than 3.0 times as long as wide …………………………………………………………………37

37. F2 longer than F3 (Fig. 2.7); radial cell 2.7 times as long as wide (Fig. 4.6) ………… ………………………………………………………………… Alloxysta brachycera Hellén, 1963 – F2 shorter than or subequal to F3; radial cell shorter or longer than 2.7 times as long as wide …38

38. F2–F4 subequal in length (Fig. 2.15); radial cell 2.8 times as long as wide (Fig. 4.13) …………… …………………………………………………………………… Alloxysta crassa ( Cameron, 1889) – F2–F4 not subequal in length; radial cell shorter than 2.8 times as long as wide …………………39

39. F1 4.4 times as long as wide; F3 longer than F4 (Fig. 2.30); radial cell 2.9 times as long as wide, Rs and R1 reaching the costal margin (Fig. 5.2) ………… Alloxysta nigrita ( Thomson, 1862)

– F1 1.4 times as long as wide, F3 subequal to F4 (Fig. 3.2); radial cell 2.8 times as long as wide; Rs and R1 not reaching the costal margin (Fig. 5.6) … Alloxysta piceomaculata ( Cameron, 1883)

40. Female: rhinaria and club shape begin on F4; F1 shorter than pedicel and slightly longer than F2; F2 subequal to F3; F3 shorter than F4 (Fig. 12.3); radial cell 2.0 times as long as wide (Fig. 12.1). Male unknown ………………………………………… Alloxysta pascuali Ferrer-Suay sp. nov.

– Specimens without the same combination of features …………………………………………41

41. Female: rhinaria and club shape begin on F3; F1 2.9 times as long as wide; F2 shorter than F3; F3 subequal to F4 (Fig. 3.19). Male unknown … Alloxysta xanthopa ( Thomson, 1862) – Rhinaria and club shape begin on F2, different proportion between flagellomeres ………42

42. Female: F1 5.2 times as long as wide; F2–F4 subequal in length (Fig. 2.32). Male: F2 slightly longer than F3; propodeum with two carinae well defined and separated on the anterior half with setae present, joining forming a plate in the posterior half; radial cell 2.6 times as long as wide (Fig. 5.4) …… Alloxysta pallidicornis ( Curtis, 1838)

– Female: F1 4.1 times as long as wide; F2 longer than F3; F3 longer than F4 (Fig. 2.1). Male: F2 shorter than F3; propodeum with two carinae forming a thick plate, with setae on top and curved sides; radial cell 2.2 times as long as wide (Fig. 4.1) … Alloxysta abdera Fergusson, 1986

43. Propodeal carinae absent …………………………………………………………………………44 – Propodeal carinae present …………………………………………………………………………50

44. Pronotal carinae absent ……………………………………………………………………………45 – Pronotal carinae present …………………………………………………………………………46

45. F1 longer than pedicel, F1 3.0 times as long as wide (Fig. 3.15); radial cell 2.3 times as long as wide; Rs reaching the costal margin (Fig. 5.19) …………………… Alloxysta soluta Hellén, 1963

– F1 subequal to pedicel and longer than F2; F2 subequal to F3 (Fig. 2.33); radial cell 2.8 times as long as wide, Rs not reaching the costal margin (Fig. 5.5) … Alloxysta patens Hellén, 1963

46. Rhinaria and club shape begin on F4 ……………………………………………………………47 – Rhinaria and club shape begin on F3 ……………………………………………………………49

47. F2–F4 subequal in length …………………………………… Alloxysta fuscipes ( Thomson, 1862) – F2–F4 not subequal in length ……………………………………………………………………48

48. Female: F1 subequal to F2; F2 longer than F3; F3 subequal to F4 (Fig. 4.21). Male: F2 and F3 curved; F1 subequal to F2; F3 longer than F4; radial cell very big, 3.0 times as long as wide in both male and female (Fig. 8.6) ……… Alloxysta macrophadna ( Hartig, 1841)

– Female: F1 longer than F2; F2 shorter or subequal to F3; F3 shorter than F4 (Fig. 2.31). Male: without any flagellomere curved; F1 longer than F2; F3 shorter than F4; radial cell 2.7 times as long as wide in both male and female (Fig. 5.3) …… Alloxysta obscurata ( Hartig, 1840)

49. F1 6.3 times as long as wide, F2 4.6 times as long as wide (Fig. 3.11); radial cell 2.6 times as long as wide (Fig. 5.15). Male unknown ……………………… Alloxysta salicicola Belizin, 1973

– F1 3.5 times as long as wide; F2 2.1 times as long as wide (Fig. 3.13); radial cell 2.7 times as long as wide (Fig. 5.17). Male: rhinaria and club shape begin on F4; F1 longer than F2; F2 longer than F3; F3 shorter than F4 ………………………………… Alloxysta semiaperta Fergusson, 1986

50. Propodeal carinae not protruding; F1 subequal to pedicel in both sexes; rhinaria and club shape begin on F4 (Fig. 2.13);radial cell 2.1 times as long as wide (Fig. 4.11) … Alloxysta citripes ( Thomson, 1862) – Propodeal carinae well defined and protruding; without the same combination of features …51

51. Propodeum with two carinae well defined, reaching the posterior part of the propodeum independently, thick and with the sides curved; rhinaria and club shape begin on F 3 in female (Fig. 3.4); F1–F3 slightly curved in male; radial cell small with Rs vein straight (Fig. 5.8) ………… ………………………………………………………………… Alloxysta pleuralis ( Cameron, 1879)

– Propodeum with two carinae forming a plate or only joining at the base; without the same combination of features ……………………………………………………………………………52

52. Rhinaria and club shape begin on F4 ……………………………………………………………53 – Rhinaria and club shape begin on F2 or F3 ……………………………………………………57

53. Pronotal carinae absent ……………………………………………………………………………54 – Pronotal carinae present ……………………………………………………………………………55

54. Female: pedicel-F3 subequal in length (Fig. 3.10); propodeum with two carinae forming a plate with straight sides …………………………………………… Alloxysta rufiventris ( Hartig, 1840)

– Female: pedicel-F3 not subequal in length (Fig. 3.14); propodeum with two carinae forming a plate with only slightly curved sides …… Alloxysta slovenica Ferrer-Suay & Pujade-Villar, 2013

55. F1 subequal to pedicel ………………………………………… Alloxysta postica ( Hartig, 1841) – F1 longer than pedicel ……………………………………………………………………………56

56. Female: F1 4.6 times as long as wide; F2 3.3 times as long as wide; F3 3.1 times as long as wide; F2 subequal to F3 (Fig. 2.11); radial cell 2.3 times as long as wide (sometimes the club shape begins on F3) (Fig. 4.9) ………………………………… Alloxysta castanea ( Hartig, 1841)

– F1 5.7 times as long as wide; F2 5.0 times as long as wide; F3 5.7 times as long as wide; F2 shorter than F3 (Fig. 2.5); radial cell 3.0 times as long as wide (Fig. 4.4). Male unknown…… …………………………………………………………………… Alloxysta aurata Belizin, 1968

57. Female: pedicel-F3 subequal in length (Fig. 2.27). Male: F1 longer than pedicel and F2; F2 subequal in F3; radial cell 2.0 times as long as wide (Fig. 4.23) … Alloxysta melanogaster ( Hartig, 1840)

– Female: F1 longer than pedicel and F2; radial cell 2.6 times as long as wide. Male unknown ………………………………………………………………… Alloxysta longipennis ( Hartig, 1841)

58. Propodeal carinae present …………………………………………………………………………59 – Propodeal carinae absent …………………………………………………………………………70

59. Propodeal carinae independent, slightly fused posteriorly ………………………………………… …………………………………………… Alloxysta barbotini Ferrer-Suay & Pujade-Villar, 2016 – Propodeal carinae fused forming a plate …………………………………………………………60

60. Pronotal carinae present …………………………………………………………………………61 – Pronotal carinae absent …………………………………………………………………………76

61. Female unknown. Male: radial cell 1.8 times as long as wide; club shape begins on F2 and rhinaria on F3; F1 longer than pedicel and F2; F2 longer than F3 (Fig. 2.10); all flagellomeres straight; propodeal carinae with curved sides ………………………… Alloxysta brevitarsis ( Thomson, 1862)

– Radial cell longer than 1.8 times as long as wide; without the same combination of features …62

62. Rhinaria and club shape begin on different flagellomeres; club shape begins on F2 and rhinaria on F1: F2 subequal to F3 (Fig. 3.18). Male unknown …… Alloxysta xanthocera ( Thomson, 1862) – Rhinaria and club shape begin on the same flagellomeres ………………………………………63

63. F1 longer than pedicel ……………………………………………………………………………64 – F1 shorter than or subequal to pedicel ……………………………………………………………66

64. Female: rhinaria and club shape begin on F4; radial cell 2.7 times as long as wide (Fig. 5.1). Male unknown. Propodeal carinae forming a complete plate with few setae anteriorly and with divergent peaks posteriorly …………………………………… Alloxysta nigricans Hellén, 1963

– Female: rhinaria and club shape begin on F3; with different combination of features …65

65. Female: F2 shorter than F3 (Fig. 3.7). Male: rhinaria begin on F1; pedicel–F3 subequal and slightly curved; radial cell 2.7 times as long as wide in female (Fig. 5.11) and 2.4 times in male …………………………………………………………………… Alloxysta pusilla (Kieffer, 1902)

– Female: F2 subequal to F3 (Fig. 3.3). Male: rhinaria begin on F2; pedicel–F3 not subequal, without any flagellomere curved; radial cell 2.4 times as long as wide (Fig. 5.7) ……… ………………………………………………………………… Alloxysta pilipennis ( Hartig, 1840)

66. F1 shorter than pedicel ………………………………………… Alloxysta heptatoma Hellén, 1963 – F1 subequal to pedicel ……………………………………………………………………………67

67. Rhinaria and club shape begin on F5; propodeal carinae narrow and well defined in the anterior half, wide and forming a plate in posterior half with sharp margins ……………………………… ……………………………………………………………… Alloxysta sawoniewiczi Kierych, 1988

– Rhinaria and club shape begin on F3 or F4; propodeal carinae form a complete plate with curved or slightly curved margins ………………………………………………………………68

68. Rhinaria and club shape begin on F4; F1 longer than F2; F2 subequal to F3; with two small carinae on the pronotum, sometimes difficult to see (Fig. 3.9); propodeum with two carinae separed by setae on the anterior third and forming a plate in the posterior two thirds; radial cell 2.0 times as long as wide (Fig. 5.13) …… Alloxysta ramulifera ( Thomson, 1862)

– Rhinaria and club shape begin on F3; without the same combination of features ……69

69. Female: F3 shorter than F4 (Fig. 2.4). Male: F1 longer than pedicel and subequal to F2; F2 slightly curved and shorter than F3; radial cell 2.3 times as long as wide (Fig. 4.3) ……………… …………………………………………………………………… Alloxysta arcuata (Kieffer, 1902)

– Female: F3–F4 subequal in length (Fig. 2.16); radial cell 2.6 times as long as wide (Fig. 4.14). Male unknown …………………………………………… Alloxysta crassicornis ( Thomson, 1862)

70. Head yellow; F1 longer than F2, F2–F4 subequal (Fig. 3.17); radial cell 3.0 times as long as wide (Fig. 5.21); propodeum without setae where they are usually present in other species (Fig. 9.2) …… ………………………………………………………………… Alloxysta victrix ( Westwood, 1833)

– Head brown; without the same combination of features ………………………………………71

71. Pronotal carinae absent ……………………………………………………………………………72 – Pronotal carinae present …………………………………………………………………………74

72. F1–F3 not subequal in length ………… Alloxysta kovilovica Ferrer-Suay & Pujade-Villar, 2013 – F1–F3 subequal in length …………………………………………………………………………73

73. Female: rhinaria and club shape begin on F4; F1 longer than pedicel (Fig. 2.2); radial cell 2.4 times as long as wide (Fig. 4.2). Male with antenna unknown … Alloxysta aperta ( Hartig, 1841)

– Female unknown. Male: rhinaria and club shape begin on F3; F1 subequal to pedicel (Fig. 3.8); radial cell 2.5 times as long as wide (Fig. 5.12) ……… Alloxysta quedenfeldti ( Kieffer, 1909)

74. Female: rhinaria and club shape begin on F3; F1 longer than pedicel and subequal to F2; F2 shorter than or subequal to F3 (Fig. 2.12). Male: F1–F3 not curved ……………………… ……………………………………………………………… Alloxysta circumscripta ( Hartig, 1841)

– Rhinaria and club shape begin on other flagellomere; without the same combination of features …75

75. Female: rhinaria and club shape begin on F3 or F4; F1 longer than pedicel and F2; F2 subequal to F3 (Fig. 2.14). Male: F1–F3 curved; radial cell 2.7 times as long as wide (Fig. 4.12) ………… …………………………………………………………… Alloxysta consobrina ( Zetterstedt, 1838) …………………………………………………………………… Alloxysta tscheki ( Giraud, 1860)

– Female: rhinaria and club shape begin on F2; F1 longer than pedicel and F2; F2 shorter than F3 (Fig. 2.23). Male: F1 curved; radial cell 2.0 times as long as wide (Fig. 4.19) ……………… …………………………………………………………………… Alloxysta leunisii ( Hartig, 1841)

76. F1 longer than pedicel …………………………………………… A. fracticornis ( Thomson, 1862) – F1 shorter than or subequal to pedicel ……………………………………………………………77

77. Male and female: F1 subequal to pedicel and longer than F2; F2 subequal to F3 (Fig. 2.28) … ……………………………………………………………… Alloxysta mullensis ( Cameron, 1883) – F1 shorter than pedicel, F1–F3 subequal in length, sometimes F1 slightly longer ………………78

78. Antenna shorter than body length; fore wing with normal marginal setae (Fig. 10.2) …………… …………………………………………………………………… Alloxysta brevis ( Thomson, 1862)

– Antenna subequal or longer than body length; fore wing with long marginal setae (Fig. 10.3) … ……………………………………………………………………… Alloxysta darci ( Girault, 1933)