Alloxysta macrophadna ( Hartig, 1841 )

Ferrer-Suay, Mar, Selfa, Jesús & Pujade-Villar, Juli, 2018, Palaearctic species of Charipinae (Hymenoptera, Figitidae): two new species, synthesis and identification key, European Journal of Taxonomy 427, pp. 1-110 : 48-50

Alloxysta macrophadna ( Hartig, 1841)

Figs 2.25, 4.21

Xystus macrophadnus Hartig, 1841: 352 . Type: deposited in ZSM (examined).

Allotria macrophadna – Giraud 1860: 130.

Alloxysta macrophadna – Förster 1869: 340.

Allotria macrophadnus – Cameron 1889: 55.

Dilyta macrophadnus – Kieffer 1900: 114.

Alloxysta (Alloxysta) macrophadna – Dalla Torre & Kieffer 1902: 38.

Alloxysta macrophadnus – Rohwer & Fagan 1919: 340.

Charips macrophadnus – Muesebeck & Krombein 1951: 607.

Alloxysta macrophadna – Andrews 1978: 85.

Diagnosis

Alloxysta macrophadna is mainly characterized by a big, partially open radial cell (Fig. 4.21); the presence of pronotal carinae; the absence of propodeal carinae; and antenna with rhinaria and club shape beginning on F 4 in females and F 3 in males. It is similar to A. obscurata , but can be differentiated by the proportions of the flagellomere (F1 is subequal to F2, F2 is longer than F3, and F3 is subequal to F 4 in A. macrophadna (Fig. 2.25), while F1 is longer than F2, F2 is shorter than F3, and F3 is shorter than F 4 in A. obscurata (Fig. 2.31)), flagellomere shape (F2 and F3 are strongly curved in A. macrophadna males, but no flagellomeres are curved in A. obscurata males), and the size of the radial cell (3.0 times as long as wide in A. macrophadna (Fig. 4.21) but 2.7 times in A. obscurata (Fig. 5.3)).

Material examined

Lectotype

GERMANY: ♀ [♀], [lectotype H.H. Evenhuis (orange label)], [ Xystus macrophadnus Hartig det. H.H. Evenhuis 1974], [ Alloxysta macrophadna ( Hartig, 1841) ♀ M. Ferrer-Suay det. 2011] ( ZSM).

Paralectotypes

GERMANY: 3 ♀♀ [In collection Hartig as Xystus macrophadnus ; Paralectotype Xystus macrophadnus Hartig, 1841 ♀ (red label); Alloxysta macrophadna ( Hartig, 1841) ♀ M. Ferrer-Suay det. 2011] ( ZSM). Additional specimens (33 ♂♂, 39 ♀♀)

CZECH REPUBLIC: 1 ♀ [Moravia, Zidlochovice, 20 Nov. 84, N.D. Springate] ( BMNH); 1 ♀ [central Bohemia, 30 Jul. 1991, J. Macek] ( CNCI C-278); 1 ♂ [Moravia, Lanzhot-Ranspurk, 9–12 Aug. 1991, L. Masner, climax hardwood forest, PT] ( CNCI C-270); 1 ♀ [Moravia, Ladnice environs 7–9 Aug. 1991, L. Masner riparian, forest] ( CNCI C-272).

FINLAND: 2 ♀♀ [Helsingfords, 19 Sep. 1933, G.J. Kerrich] ( BMNH B.M. 1969-261); 1 ♀ [Kevo, Res. Station, 69°45′ N, 27°00′ E, 24 Jun. 1989, H. Goulet, sweep] ( CNCI C-309).

FRANCE: 1 ♂ [Haute Savoie, Bossy sur Frangy, 17–30 Aug. 1993, J. Steffen, on wood pile] ( CNCI C-289).

GERMANY: 1 ♀ [Wern, Riedenerwald, 15 Aug. 1978, L. Huggert, sweep in open oak wood] ( CNCI C-234); 1 ♂, 5 ♀♀ [Schwarzwald, Todtmoos environs, 6 Aug. 1984, L. Masner, screen sweeping] ( CNCI C-235).

IRELAND: 1 ♀ [Co. Donegal: Saggart A.W. Stelfox, 29 Sep. 1935 (green label), Saggar (n. Dv. Aws, 29.9.35) (handwritten, green label), A.W. Stelfox Collection 1966] ( USNM).

ITALY: 1 ♂ [Dolomites Seis am Schlern., 1–13 Jun. 1964] ( BMNH).

JAPAN: 1 ♂, 2 ♀♀ [Hokkaido, Aizan, 800 m, 4 Jul. 1989, sweep, M.J. Sharkey] ( CNCI C-186); 1 ♀ [Hokkaido, Furano, Exp. Forest, 43°15′ N, 142°20′ E, 9 Aug. 1996, 500 m, L. Masner, sweep] ( CNCI C-166); 4 ♂♂, 3 ♀♀ [Hokkaido, Furano Exp. Forest, 43°15′ N, 142°20′ E, 9 Aug. 1996, 500 m, L. Masner, sweep] ( CNCI C-162); 1 ♀ [Hokkaido, Sapporo, Jozankei, 21–28 Sep. 1989, K. Maeto and M. Sharkey] ( CNCI C-193); 4 ♂♂ [Gifu, Mt. Ena-san (ssw), 1200–1900 m, 5 Jun. 1995, K. Yamagishi] ( CNCI C-189); 6 ♂♂ [Hokkaido, Shibetsu-Rubesu, E. slope of Mt. Shari, 43°45′ N, 144°45′ E, 17 Aug. 1996, 700 m, L. Masner, sweep] ( CNCI C-164); 3 ♂♂, 1 ♀ [Hokkaido, Furano, Exp. Forest, 43°15′ N, 142°20′ E, 9 Aug. 1996, 500 m, L. Masner, sweep] ( CNCI C-163).

NORWAY: 1 ♀ [Trondhem, 3 Jul. 1980, F. Joldsa] ( BMNH).

POLAND: 1 ♀ [Przemysl, 12 Sep. 1984, N.D. Springate] ( BMNH).

RUSSIA: 1 ♀ [Magadanskaya Oblast, Aborigen Field Station, 500 m, 25 Jul.–10 Aug. 1990, D.M. Wood] ( CNCI C-298).

SLOVAKIA: 1 ♀ [Koṧice, 14–17 Sep. 84, N.D. Springate] ( BMNH).

SLOVENIA: 1 ♀ [Slov. Vrata Valley, 13 Jun. 1979, P.H. and S.L. Ward] BMNH B.M. 1979-263); 1 ♂ [Rupa, 28 Apr. 1975, J.S.N., Grassland] ( BMNH); 1 ♂ [Slovenia, Slatna, Radovljica, 6 Aug. 1978, L. Huggert, swept on open wooded (deciduous) meadow] ( CNCI C-275).

SWEDEN: 1 ♂, 8 ♀♀ [Sk. Lund, Zoological Mus. grounds, Aug. 1976] ( BMNH); 1 ♂, 2 ♀♀ [Sk. Röstanga, 6 Jul. 1938, D.M.S.P. and J.F.P.] ( BMNH B.M. 1938-414); 1 ♀ [Kalmar, Överum, 19 May–29 Jun. 1978, L. Huggert, MT at swampy lake shore] ( CNCI C-312); 1 ♀ [Värmland, Ekshärad, 20–29 Jul. 1960, W.R.M. Mason] ( CNCI C-313).

SWITZERLAND: 7 ♂♂, 1 ♀ [Dielsdorf, 650 m, 17 Aug. 1984, L. Masner, sweep] ( CNCI C-290).

UNITED KINGDOM: Scotland: 1 ♀ [Perth and Kinross, Acham, PM, A.W. Stelfox, 15 Jul. 1951, A.W. Stelfox Collection 1966] ( USNM).

Distribution

Holarctic.

Certain records: Andorra ( Ferrer-Suay et al. 2011: 357), England ( Müller et al. 1999: 346), Germany ( Hartig 1841: 352; Höller et al. 1993: 13), Iran ( Ferrer-Suay et al. 2013a: 38), Montenegro ( Ferrer-Suay et al. 2013b: 358);

Uncertain records: Austria ( Giraud 1860: 130; Hellén 1963: 12), Belgium ( Lameere 1907; Crèvecoeur & Maréchal 1933), Bulgaria and Balkan peninsula ( Vasileva-Sumnalieva 1976: 23), England ( Cameron 1889: 53; Kieffer 1902a: 10), Finland ( Hellén 1963: 12), France ( Kieffer 1902a: 10; De Gaulle 1908: 26), Ireland ( O’Connor & Nash 1997), Italy ( Hellén 1963: 12), Lappland ( Hellén 1963: 12), Norway ( Hellén 1966: 393), Poland ( Kierych 1979b: 14; Krawczyk et al. 2009: 161), Romania ( Barnea et al. 2005: 87), Russia ( Hellén 1963: 12), Scotland ( Cameron 1883: 368; 1886: 53), Sweden ( Thomson 1862: 408), Switzerland ( Hellén 1963: 12), The Netherlands ( Evenhuis 1974: 165), USA (Tennessee) ( Andrews 1978: 59).

New records: Czech Republic and Japan.