Laneella, Mello, 1967

Whitworth, Terry L. & Yusseff-Vanegas, Sohath, 2019, A revision of the genera and species of the Neotropical family Mesembrinellidae (Diptera: Oestroidea), Zootaxa 4659 (1), pp. 1-146 : 24

Key to species of Laneella

1 Basal half of tarsal claws pale, white to pale yellowish ( Fig. 10 ); wing with infuscation along costa or tip of wing; male with basal 2/3 of cerci slender in dorsal view, as in Figs 20 , 22 . Female terminalia as in Figs 284–285...................... 2

- Basal half of tarsal claw orange, brown or black ( Fig. 11 ); area along costa with at most faint infuscation, distal portion of wing not infuscated; male with basal 2/3 of cerci much broader in dorsal view in two of three species ( Figs 18 , 24 ). Female terminalia distinctive, as in Figs 282–283, 286....................................................................... 3

2 Wing with anterior edge infuscated from junction of subcosta-costa posteriorly to R 4+5 and distally to tip; palpus brown to black; row of black postocular setae ending about 2/3 of way from top of eye to gena, with pale yellowish below that; anterior half of T4 with solid whitish band of tomentum; fore and mid femora dark reddish-brown, hind femur orange to brown; [known from Brazil and Paraguay].......................................................................... L. nigripes

- Wing with distal 1/2–1/3 of tip infuscated, also cells r 1, r 2+3 and r 4+5 and junction of vein CuA1 and crossvein dm-cu darkened; palpus orange; row of black postocular setae reaching gena; midsegment of T4 with four transverse pale spots; all femora orange; [known from Brazil, Colombia, Costa Rica and Ecuador]........................................... L. perisi

3 Thorax and abdomen dark metallic bluish-purple ( Fig. 457 ); base of wing with cells and veins darkened; disc, rims and setae of calypters dark brown; anterior half of basicosta brown, posterior half more or less reddish-brown; [known primarily from western and central Costa Rica].......................................................... L. purpurea sp. nov.

- Thorax brown to reddish-brown, not metallic; base of wing not darkened; calypters with disc, rims and fringe of setae light to moderately brown; basicosta orange...................................................................... 4

4 Wing with distinctive brown spot around crossvein r-m ( Fig. 3 ); abdomen shiny brown or bluish-purple; wing with section IV about 0.25 of section III or less.......................................................................... 5

- Wing without darkened area around crossvein r-m; abdomen shiny metallic blue to purple; wing with section IV about 0.46 of section III; [known from Mexico and Guatemala]........................................ L. fuscosquamata sp. nov.

5 Abdomen bluish-purple, mid and hind femora with apices dark brown; [known only from Colombia]........... L. patriciae

- Abdomen shiny brown; mid and hind femora with apices orange; [known from Costa Rica, Ecuador and Venezuela].......................................................................................... L. fusconitida sp. nov.

Figures

FIGURES 9–14. Diagnostic characters of Mesembrinellidae. 9. Eggs of Laneella fuscosquamata sp. nov. 10. L. perisi (Mariluis), ventral view of tarsal claws, showing pale basal half. 11. L. fusconitida sp. nov., ventral view of tarsal claws, showing darker base. 12. M. spicata Aldrich, ventral view of male sternites, showing long projections on rear margin of ST5. 13. M. bicolor (Fabricius), posterior view of female occiput, showing median occipital sclerite. 14. M. socors (Walker), upper and lower calypters, showing disc, rim and fringe.

FIGURES 15–20. Left lateral and posterior views of epandrium, cerci and surstyli of Laneella spp. 15–16. L. fusconitida sp. nov. 17–18. L. fuscosquamata sp. nov. 19–20. L. nigripes Guimar„es.

FIGURES 21–26. Left lateral and posterior views of epandrium, cerci and surstyli of Laneella and Mesembrinella spp. 21–22. L. perisi (Mariluis). 23–24. L. purpurea sp. nov. 25–26. M. latifrons (Mello).

FIGURES 282–287. Female terminalia (T6–T8, epiproct, cerci) of Mesembrinellidae, dorsal view. 282. Laneella fusconitida sp. nov. 283. L. fuscosquamata sp. nov. 284. L. nigripes Guimar„es. 285. L. perisi (Mariluis). 286. L. purpurea sp. nov. 287. M. latifrons (Mello).

FIGURES 455–459. Lateral habitus and labels of holotypes. 455. Laneella fusconitida sp. nov. (LACM). 456. L. fuscosquamata sp. nov. (USNM). 457. L. purpurea sp. nov. (INBIO). 458. Mesembrinella mexicana sp. nov. (CNC). 459. M. spicata Aldrich (USNM).

FIGURES 1–8. Diagnostic characters of Mesembrinellidae and Calliphoridae. 1. Wing of Mesembrinella flavicrura Aldrich, showing veins and cells. 2. Wing of M. currani Guimar„es, showing infuscated anterior margin. 3. Wing of Laneella fusconitida sp. nov., showing dark macula around crossvein r-m. 4. Wing of Calliphora vomitoria (Linnaeus), showing acute bend of vein M. 5. Anterior spiracle of Mesembrinella batesi Aldrich, showing broad oval opening above. 6. Anterior spiracle of M. bolivar (Bonatto), showing long, narrow opening gradually widening above. 7. Meral setae and posterior spiracle of M. bicolor (Fabricius). 8. Last abdominal tergites of M. vogelsangi (Mello), showing setae.