Eupsenella kaibara Ramos & Azevedo

Ramos, Magno S. & Azevedo, Celso O., 2012, Revision of Eupsenella Westwood, 1874 (Hymenoptera, Bethylidae), Zootaxa 3539 (1), pp. 1-80 : 47-48

Eupsenella kaibara Ramos & Azevedo , sp. nov.

( Figs 36 , 81, 126)

Description. FEMALE (holotype). Body length 8.67 mm. LFW 3.81 mm.

Coloration. Head dark castaneous almost black; scape dark castaneous; pedicel and flagellum castaneous; mandible dark castaneous; palpi light castaneous; mesosoma dark castaneous almost black; tegula dark castaneous; wing venation dark castaneous; procoxa dark castaneous almost black; mesocoxa and metacoxa dark castaneous; trochanters light castaneous; profemur dark castaneous almost black; mesofemur, metafemur and tarsi castaneous; metasoma dark castaneous almost black.

Head ( Fig. 36 ). Head as long as wide, globoid in profile; gena strongly concave in profile. Mandible evenly wide, two lowermost teeth larger than upper ones, lower tooth curved inward, lower margin strongly setose. Clypeus coriaceous, median clypeal lobe angulate, median concavity of median clypeal lobe absent, 0.07 LH, well marked off from frons, median carina present, 0.15 LH. Malar space conspicuous, 0.38 mandibular base. Antenna 1.90 mm; first five antennomeres in ratio of about 14:13:13:14:14; flagellomere 1 1.3 longer than wide, flagellar pubescence dense, with few outstanding erect setae. Region between torulus and eye straight. Area between the anterior margin of eyes and demarkation of median clypeal lobe 0.08 LH. Eye protuberant, 0.54 LH, with sparse hairs, located more laterally, lower margin without strong depression. Frons coriaceous-punctate. WH 1.0 LH. WF 0.59 WH. WF 1.07 HE. OOL 1.0 WOT. VOL 0.33 HE. Distance from posterior ocellus to vertex crest 0.67 DAO. Ocelli large, DAO 0.12 WF, ocellar triangle not compact, distance between posterior ocelli 2.0 DAO, anterior ocellus surpassing imaginary top eye line, posterior ocellus almost reaching vertex. Vertex straight, short setae, 0.15 LH, temple profile slightly convergent in dorsal view, with corner angulate, not projected postero-laterad. Hipostomal carina sinuous, low.

Mesosoma (Fig. 81). Pronotal disc coriaceous-punctate, 0.24 as long as wide, anterior corner angulate, coriaceous-punctate, 0.75 length pronotal disc, almost flat in profile, posterior margin almost straight medially, scutellar pit elliptical, dilated, distant each other 8.67 its own diameter. Dorsal axillar surface with fovea oval. Metanotum with median region almost flat, lower than scutellum in profile, with fovea oval laterally. Propodeal disc wider than long; anterior margin without foveolate or transverse furrow; median carina 0.88 length propodeal disc, incomplete posterad; discal carina 0.13 length propodeal disc, converging posterad, slightly arched; space between median and discal carina confused-rugulose. Mesopleuron with subtegular furrow oval; mesepimeral suture almost inconspicuous posterad, mesopleural epicoxal lobe inconspicuous. Propleuron with sequence of foveae from anterior margin of prosternum until cervical collar absent. Prosternum with median excavation complete, median suture present. Mesopectus with posterior depression heart-shaped; median suture evident.

Wings. Macropterous. Forewing ( Fig. 126 ) with 1M cell subpentagonal; 1M and 1R1 cells not aligned basally; C cell widening distally;1R1 cell present, slightly longer than 2R1 cell, evenly wide; Rs b convex; 2R1 cell 0.16 LFW; distal margin of Rs c convex; r-rs vein inclined; R1 b entirely on anterior margin of forewing; Cu b absent; Rs+M vein 1.45 Rs a; M vein 1.67 Rs a; Rs a subperpendicular to anterior margin of forewing; stigma rectangular; distal stigmal margin truncate. Hind wing with four distal hamuli, last hamulus discontinuous each other.

Legs. Profemur swollen, 0.55 as wide as long. Metacoxa with median region 0.59 as wide as long. Metacoxal projection in dorsal region conspicuous, apex rounded.

Metasoma. 0.2 as wide as long, ventral carina of petiole present.

MALE. Unknown.

Material examined. Holotype ♀. AUSTRALIA, 35.22S 148.50E, A[ustralian] C[apital] T[erritory], Blundells C [ree]k, Feb [ruary] 1987, D.H.Colless, Malaise trap /ethanol, ANIC . Paratypes. AUSTRALIA, W[estern] A[ustralia], 1 ♀, Crowea S [ta]t[e] For [est], n[ea]r Pemberton, 29 Oct [ober]–20 Dec [ember] 1979, S.J.Curry, Malaise trap, closed forest, ANIC ; A[ustralian] C[apital] T[erritory], 1 ♀, 35.22S 148.50E, Blundells C [ree]k, Feb [ruary] 1987, D.H.Colless, Malaise trap /ethanol, ANIC .

Distribution. Australia (Australian Capital Territory, Western Australia).

Variation. Body length 4.76–5.25 mm; LFW 3.80–4.04 mm; median carina of clypeus high; veins of forewing light castaneous.

Remarks. This species is similar to E. barada sp. nov. by having the head subtriangular in profile, the eyes protuberant, densely hairy. However this species has the area between the anterior margin of eyes and demarkation of median clypeal lobe short, the body robust, the median clypeal lobe with high median carina, the gena slightly concave in profile, the vertex crest slightly convex, the notauli parallel, whereas E. barada sp. nov. has the area between the anterior margin of eyes and demarkation of median clypeal lobe long, the median clypeal lobe truncate, without median carina, the vertex crest slightly concave, the notauli convergent posterad.

Figures

FIGURES 27–38. Head in dorsal view. 27. Eupsenella ingura sp. nov. 28. E. iwaidja sp. nov. 29. E. jaadwa sp. nov. 30. E. jaara sp. nov. 31. E. jaburara sp. nov. 32. E. jadira sp. nov. 33. E. jagara sp. nov. 34. E. janda sp. nov. 35. E. kabalbara sp. nov. 36. E. kaibara sp. nov. 37. E. karanja sp. nov. 38. E. karawa sp. nov. (Scale bar = 300 µm).

FIGURES 123–132. Right forewing. 123. Eupsenella jagara sp. nov. 124. E. janda sp. nov. 125. E. kabalbara sp. nov. 126. E.