Quadraticossus fangi, Ying, Wang & Dong, Ren, 2007

Ying, Wang & Dong, Ren, 2007, Two new genera of fossil palaeontinids from the Middle Jurassic in Daohugou, Inner Mongolia, China (Hemiptera, Palaeontinidae), Zootaxa 1390, pp. 41-49 : 43

Quadraticossus fangi sp. nov.

( Fig. 1A–F )

Diagnosis: Sc without branches and terminating on R1 near the distal part. M bifurcated into M1+2 and M3+4 at level of Rs arising from R. Horizontal crossvein m4­cua long.

Description: Head with pair of large eyes. Rostrum long but not reaching posterior margin of mesoscutum. In ventral view, legs partly preserved, femora stout, tibiae somewhat slender. Abdomen with at least 6 visible segments.

Forewing triangular with distinct CP and nodal line. Sc without branches, separated from R at base, then fused with R after point of R separating into R1 and Rs, extending beyond nodal line and terminating on R1. M bifurcated into M1+2 and M3+4 at same level of Rs arising from R. M1+2 and M3+4 separated at same level. Crossvein r­m present between Rs and M1. Cu divided into CuA and CuP at base. CuA curved, irregular, joined with M by crossvein m­cua and also connected with M4 by long horizontal crossvein. CuA branched into CuA1 and CuA2 after point of M bifurcating into M1+2 and M3+4. CuP single. Vein A two branched, A2 with branches. Nodal line traceable as indentation across Sc, and along with Sc for a short distance, then across Rs to M1+2, along with the latter to branching point of M, separating discal cells into two unequal parts, following stem CuA and CuA2, terminating at distal end of clavus.

Hind wing smaller than forewing, costal margin curved posteriorly, forming prominent indentation. Rs coalesced with M1 for long interval before separating. Stem of M1+2 long, M2 arising from M1 at same level of Rs separating from R. M4 not divided from M3+4. Short crossvein m3+4­cua present. CuA1 fused with CuA2 for long interval. Anal vein single. Both forewings and hind wings with prominent color pattern consisting of hyaline patches on a dark ground. ( Fig.1. A, B, C , D, F ).

Body 25 mm long, 15 mm wide; forewing 41mm long, 18 mm wide; hind wing 24 mm long, 16 mm wide.

Etymology: This species is named after Mr. Fang Liang for his contribution in collecting Daohugou fossils, including this specimen.

Material: The new species is established based on body with a pair of forewings and a pair of hind wings. Holotype: Body with wings compression, part and counterpart, No.: CNU­H­NN2006093­1, CNU­H­ NN2006093­2.

Horizon and locality: Jiulongshan Formation, Middle Jurassic, Daohugou Village, Shantou Township, Ningcheng County, Inner Mongolia, China.

Figures

FIGURE 1 A – C. Quadraticossus fangi gen. & sp. nov., holotype: A, photograph of part CNU­H­NN 2006093 ­ 1; B, photograph of counterpart CNU­H­NN 2006093 ­ 2; C, photograph of subcosta (Sc) on forewing.

FIGURE 1 D – F. Quadraticossus fangi gen. & sp. nov., holotype: D, body with wings; E, forewing; F, hind wing.