Sagola monticola Broun, 1912

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 136

Sagola monticola Broun, 1912

( Figs. 68h , 69h , 70h , 72 )

Sagola monticola Broun, 1912a: 402 . Hudson 1923: 365, 1934: 184; Raffray 1924: 233; Kuschel 1990: 48; Newton and Thayer 2005b; Nomura and Leschen 2006: 243.

Type Material. O n e o f s y n t y p e s. N E W ZEALAND: Taupo: 1♂ ( BMNH), glued on rectangular card, “ Type ” [red label, printed]; “3195. ♂ ” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Ngauruhoe. march. 1910.” [white label, handwritten]; “ Sagola ♂. monticola.” [white label, handwritten] .

Additional Material (n = 3: 2 males; 1 female). NEW ZEALAND: Bay of Plenty : 1♂, Rotorua, Arikikapakapa Reserve , 25 VII 1951, moss ; Taupo: 1♂ 1♀ , Desert Rd, Mangatawai , 13 I 1972, G.W. Ramsay, litter 72/1 .

Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.4–2.7 mm; temporal lobe of male head projecting forward posterio-laterally with weakly concave ventral surface; ventral surface of male head with transverse process behind mouthparts and round depression bearing three patches of dense setae; male elytra rectangular, female elytra subquadrate; shape of antennomeres and genitalia unique to species.

Redescription. Length 2.4–2.7 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler ( Fig. 68h ). Head: Male head triangular, widest across temples ( Fig. 69h ). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–7 longer than wide, 8–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round and partially covered by frontal rostrum, posterior frontal fovea round. Eye prominent, approximately one-half length of temple. Temporal lobe of male head projecting forward posterior-laterally with weakly concave ventral surface ( Fig. 69h ). Ventral surface of male head with transverse process behind mouthparts and round depression bearing 3 patches of dense setae ( Fig. 69h ). Thorax: Prosternum as long as wide, widest at one-third length. Male elytra rectangular ( Fig. 68h ), female elytra subquadrate. Male meso- and metathorax trapezoidal, longer than wide, in female as long as wide. Abdomen: Male tergite IV with pair of small transverse patches of microtrichia reaching middle, patches absent in female. Aedeagus: Median lobe of genitalia widest at midpoint and narrower apically ( Fig. 70h ). Phallobase of median lobe symmetrical and rounded ( Fig. 70h ). Parameres symmetrical and slender, with sparse setae from apex to midpoint ( Fig. 70h ).

Type Locality. Mount Ngauruhoe , Taupo, New Zealand .

Distribution. Bay of Plenty, Taupo ( Fig. 72 : triangles).

Habitat. Specimens of this species were collected by sifting moss or leaf litter.

Figures

Fig. 68. Habitus of Sagola species in the insignis species-group, dorsal views. a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 1 mm.

Fig. 69. Ventral views of male heads of Sagola species in the insignis species-group. a) Sagola insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 0.1 mm.

Fig. 70. Sagola species in the insignis species-group. Aedeagi, dorsal views: a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Male heads, dorsal surface: y) S. bipunctata, z) S. eminen. Scale bars = 0.1 mm.

Fig. 72. Known collection localities of New Zealand Sagola species. S. pertinax: black circles; S. monticola: triangles; S. laticeps: black squares; S. castanea: stars; S. sheldoni: white circles; S. mercuryensis: white squares.