Sagola angulifera Broun, 1911

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 131-132

Sagola angulifera Broun, 1911

( Figs. 68d , 69d , 70d , 71 )

Sagola angulifera Broun, 1911: 491 . Hudson 1923: 365, 1934: 183; Raffray 1924: 232; Newton and Thayer 2005b; Nomura and Leschen 2006: 241.

Type Material. Holotype. NEW ZEALAND: Taupo : 1♂ ( BMNH), glued on rectangular card, “Type” [red label, printed]; “3363.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Waimarino. Jany 1909.” [white label, handwritten]; “ Sagola ♂ angulifer” [white label, handwritten].

Additional Material (n = 19: 10 males; 9 females). NEW ZEALAND: Wellington : 2♂♂ 4♀♀, Nikau Res, Paraparaumu , 11 VI 1988, J. Nunn ; 4♂♂ 2♀♀, Nikau Res, Paraparaumu , 2 VII 1988, J. Nunn ; 2♂♂, Nikau Res, Paraparaumu , 23 VII 1988, J. Nunn ; 1♂, Nikau Res, Paraparaumu , 1 V 1988, J. Nunn, leaf litter ; 1♀, Nikau Res, Paraparaumu , 8 V 1988, J. Nunn ; 2♀♀, Nikau Res, Paraparaumu , 25 IV 1988, J. Nunn ; Taranaki: 1♂ , Potaema Walk, Mt. Egmont , 9 XII 1995, J. Nunn, forest litter .

Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.3–2.7 mm; temporal lobe of male head distinctly projecting, exceeding eye, with concave ventral surface; ventral surface of male head with transverse process behind mouthparts with three patches of long and dense setae medially; male elytra rectangular, female elytra subquadrate; shape of antennomeres and genitalia unique to species.

Redescription. Length 2.3–2.7 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler ( Fig. 68d ). Head: Male head triangular, widest across temples ( Fig. 69d ). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4–7 longer than wide, 8–10 subquadrate. Frontal sulcus deep, reaching end of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye prominent, approximately one-third length of temple. Temporal lobe of male head distinctly projecting, exceeding eye, with concave ventral surface ( Fig. 69d ). Ventral surface of male head with transverse process behind mouthparts with 3 patches of long, dense setae medially ( Fig. 69d ). Thorax: Prosternum as long as wide, widest at one-third length. Male elytra rectangular ( Fig. 68d ), female elytra subquadrate. Male meso- and metathorax trapezoidal, longer than wide, female as long as wide. Abdomen: Male tergite IV with pair of small, round patches of microtrichia, patches absent in female. Aedeagus: Median lobe of genitalia broad, with pair of triangular processes apically ( Fig. 70d ). Apical lobe with semicircular depression apically ( Fig. 70d ). Phallobase of median lobe symmetrical and rounded. Parameres symmetrical and slender, setose from midpoints to apices ( Fig. 70d ).

Type locality. Waimarino , Taupo, New Zealand .

Distribution. Taranaki, Taupo, Wellington ( Fig. 71 : stars).

Habitat. Specimens of this species were collected by sifting forest litter.

Figures

Fig. 68. Habitus of Sagola species in the insignis species-group, dorsal views. a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 1 mm.

Fig. 69. Ventral views of male heads of Sagola species in the insignis species-group. a) Sagola insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 0.1 mm.

Fig. 70. Sagola species in the insignis species-group. Aedeagi, dorsal views: a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Male heads, dorsal surface: y) S. bipunctata, z) S. eminen. Scale bars = 0.1 mm.

Fig. 71. Known collection localities of New Zealand Sagola species. S. insignis: black circles; S. hectorii: triangles; S. bipunctata: black squares; S. angulifera: stars; S. eminens: white circles; S. robustula: white squares.