Sagola hanae Park and Carlton

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 143-144

Sagola hanae Park and Carlton , new species

( Figs. 68s , 69s , 70s , 74 )

Type Material. Holotype. NEW ZEALAND: Wairarapa : 1♂ ( NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “New Zealand WA Castlehill / Haunui area. SE Puketoi Ra March 2010 ”, “ HOLOTYPE Sagola hanae Park and Carlton 2013 ” The Original label does not indicate who collected the holotype, but it was collected by J. Nunn. Paratypes (n = 6: 3 males; 3 females). NEW ZEALAND: Wairarapa : 1♂ 2♀♀ ( JTN), same data as holotype ; Wellington: 1♂ ( FMNH), Wilton’ s Bush , 110m, 41”15.963′S 175”45.159′E, 24 XI 2005, mixed broadleafpodocarp forest, FMHD#2005-030 , litter, M. Thayer, A. Newton, ANMT site 1150 ; Rangitikei: 1♂ 1♀ ( NZAC), Utiku, Main Trunk Line , J. Ford, 27 VII 1917 .

Etymology. This species is named for one of the enthusiastic supporters of this study, Hana Kim. She is one of the first author’ s colleagues and assisted with molecular components of the phylogenetic analysis.

Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.2–2.4 mm; temporal lobe of male head distinctly projecting, exceeding eye, with C-shaped ventral surface; male temporal lobe with L-shaped depression anteriorly; ventral surface of male head with transverse process behind mouthparts and round depression bearing three patches of dense setae; male elytra rectangular, female elytra subquadrate; shape of antennomeres and genitalia unique to species.

Description. Length 2.2–2.4 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler ( Fig. 68s ). Head: Male head triangular, widest across temples ( Fig. 69s ). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2 subquadrate, 3–8 longer than wide, 9–10 subquadrate. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Eye prominent, approximately one-third length of temple. Temporal lobe of male head distinctly projecting, exceeding eye, with C-shaped ventral surface ( Fig. 69s ). Male temporal lobe with L-shaped depression anteriorly ( Fig. 69s ). Ventral surface of male head with transverse process behind mouthparts and round depression bearing 3 patches of dense setae ( Fig. 69s ). Thorax: Prosternum as long as wide, widest at one-third length. Male elytra rectangular ( Fig. 68s ), female elytra subquadrate. Male meso- and metathorax trapezoidal, longer than wide, female as long as wide. Abdomen: Male tergite IV with pair of small round patches of microtrichia, patches absent in female. Aedeagus: Median lobe of genitalia widest at middle, with round apical lobe ( Fig. 70s ). Phallobase of median lobe symmetrical and rounded ( Fig. 70s ). Parameres symmetrical and triangular, setose from midpoints to apices ( Fig. 70s ).

Distribution. Rangitikei, Wairarapa, Wellington ( Fig. 74 : black circles).

Habitat. One specimen was collected by sifting forest litter from a broadleaf and podocarp forest.

Figures

Fig. 68. Habitus of Sagola species in the insignis species-group, dorsal views. a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 1 mm.

Fig. 69. Ventral views of male heads of Sagola species in the insignis species-group. a) Sagola insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 0.1 mm.

Fig. 70. Sagola species in the insignis species-group. Aedeagi, dorsal views: a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Male heads, dorsal surface: y) S. bipunctata, z) S. eminen. Scale bars = 0.1 mm.

Fig. 74. Known collection localities of New Zealand Sagola species. S. hanae: black circles; S. gyudongi: triangles; S. egmontensis: black squares; S. boonei: stars; S. stewartensis: white circles; S. jeongnamae: white squares.