Sagola buckleyi Park and Carlton

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 140-141

Sagola buckleyi Park and Carlton , new species

( Figs. 68n , 69n , 70m , 73 )

Type Material. Holotype. NEW ZEALAND: Northland: 1♂ ( NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND:ND, Poor Knights Islands Tawhiti Rahi, Main Track, XII-182009 TB270, T. R. Buckley, R. Leschen, D. Seldon sift flood debris”, “ HOLOTYPE Sagola buckleyi Park and Carlton 2013 ” . Paratypes (n = 20: 15 males; 5 females). NEW ZEALAND: Northland : 5♂♂ 4♀♀ ( DSC), same as holotype ; 3♂♂ ( NZAC), Poor Knights I, Aorangi, ridge to Oneho Hill , 17 XI 1981, J.C. Watt, litter ; 2♂♂ ( NZAC), Poor Knights I, Aorangi, Puweto Valley , 14 XI 1981, J.C. Watt, litter ; 2♂♂ ( NZAC), Poor Knights I, Tawhiti Rahi, Shag Bay , 40m, 20 IX 1980, J.C. Watt, litter ; 2♂♂ ( NZAC), Poor Knights I, Tawhiti Rahi , 4 XII 1980, G. Kuschel, litter ; 1♂ 1♀ ( NZAC), Poor Knights I, Tawhiti Rahi , Shag Bay, 30m, 12 IX 1980, J.C. Watt, litter .

Etymology. This species is named for the collector of the holotype, Thomas R. Buckley.

Diagnosis. This species can be distinguished from the other species of the insignis speciesgroup by the following combination of characters: body length 2.4–2.7 mm; temporal lobe of male head distinctly projecting, exceeding eye, with weakly concave ventral surface; ventral surface of male head with transverse process behind mouthparts and round setose depression; elytra subquadrate; shape of antennomeres and genitalia unique to species; only known from Poor Knights Islands of Northland.

Description. Length 2.4–2.7 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler ( Fig. 68n ). Head: Male head triangular, widest across temples ( Fig. 69n ). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2 longer than wide, 3 subquadrate, 4 longer than wide, 5–10 subquadrate. Frontal sulcus deep, reaching end of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea oval. Eye large and prominent, approximately one-third length of temple. Temporal lobe of male head distinctly projecting, exceeding eye and with weakly concave ventral surface ( Fig. 69n ). Ventral surface of male head with transverse process behind mouthparts and round setose depression ( Fig. 69n ). Thorax: Prosternum as long as wide, widest at one-third length. Elytra subquadrate ( Fig. 68n ). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Male tergite IV without patch of microtrichia. Aedeagus: Median lobe of genitalia broad with pair of triangular processes apically, apical lobe triangular ( Fig. 70n ). Phallobase of median lobe symmetrical and rounded ( Fig. 70n ). Parameres symmetrical and slender, with sparse setae at apex and midpoint ( Fig. 70n ).

Distribution. Only known from Poor Knights Islands, Northland ( Fig. 73 : triangle).

Habitat. Specimens of this species were collected by sifting leaf litter.

Figures

Fig. 68. Habitus of Sagola species in the insignis species-group, dorsal views. a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 1 mm.

Fig. 69. Ventral views of male heads of Sagola species in the insignis species-group. a) Sagola insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 0.1 mm.

Fig. 70. Sagola species in the insignis species-group. Aedeagi, dorsal views: a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Male heads, dorsal surface: y) S. bipunctata, z) S. eminen. Scale bars = 0.1 mm.

Fig. 73. Known collection localities of New Zealand Sagola species. S. cuvierensis: black circle; S. buckleyi: triangle; S. brumfieldi: black squares; S. otteai: stars; S. boudreauxae: white circle; S. taegyui: white squares.