Sagola hirtalis, Broun, 1893 b:, 1050

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 7-9

1. hirtalis species-group (16 species)

Diagnosis. Members of the hirtalis speciesgroup can be distinguished from other Sagola species-groups by the following combination of characters: body length 2.0–4.0 mm; gular surface of male head with deep, round depression; male antennomeres longer than those of female, antennomere 1 at least 2 times longer than wide, with dull surface; prosternum with lateral procoxal foveae; male fore femur with semicircular depression ( Fig. 4u : arrow); abdominal tergites IV–VI with discal carinae; abdominal ventrites IV–VI with basolateral foveae; parameres distinctly asymmetrical and finely punctured ( Fig. 4a–p ); present on North Island, not known from South Island ( Figs. 5–7 ).

KEY TO SPECIES OF THE HIRTALIS SPECIES- GROUP

The key is based on male specimens because most female specimens are indistinguishable based on external morphology.

1. Hind wings reduced to small pads; abdominal tergite IV without patches of microtrichia.....................................2

1′. Hind wings well-developed; abdominal tergite IV with pair of transverse patches of microtrichia reaching middle............3

2(1). Smaller body size, length 2.4 mm; antennomere 1 approximately 2 times longer than wide, 4–5 subquadrate; mid-tibia bent ( Fig. 12o : arrow) ........ ..................... S. brookesi new species

2′. Larger body size, length 2.5–2.9 mm; antennomere 1 approximately 2.5 times longer than wide, 4–5 longer than wide; mid-tibia straight ................................... .................. S. hunuaensis new species

3(1′). Larger body, length 3.2–4.0 mm; antennomere 5 elongate, at least 1.5 times longer than wide.................................... 4

3′. Smaller body, length 2.0– 3.3 mm; antennomere 5 elongate, not exceeding 1.5 times longer than wide ................... 5

4(3). Apical lobe of genitalia bird-head-shaped; left paramere with two minor slender lobes, originating from base ( Fig. 4b ) .................................. S. terricola Broun

4′. Apical lobe of genitalia shaped differently; left paramere with one lobe originating anteriorly ( Fig. 4a ).............. ................................... S. hirtalis Broun

5(3′). Smaller, body length, 2.0– 2.5 mm; antennomere 1 not exceeding 2.5 times longer than wide...............................6

5′. Larger, body length, 2.4–3.2 mm; antennomere 1 approximately 3 times longer than wide.................................... 8

6(5). Phallobase of median lobe symmetrical and elongate ( Fig. 4c ) ........................... ................................. S. convexa Broun

6′. Phallobase of median lobe asymmetrical and triangular.................................7

7(6′). Eye small, one-third length of temple; mid-femur and tibia normal; apical lobe of genitalia curved; left paramere rectangular ( Fig. 4k ).................................... ....................... S. thayerae new species

7′. Eye large, as long as temple; mid-femur with semicircular depression; mid-tibia bent; apical lobe of genitalia straight; left paramere L-shaped ( Fig. 4n ) .......... ................ S. gisbornensis new species

8(5′). Apical lobe of genitalia bulbous and semicircular..........................................9

8′. Apical lobe not bulbous and not semicircular ...............................................12

9(8). Process, either slender or triangular, extending from middle of apical lobe of genitalia .........................................10

9′. Middle of apical lobe of genitalia lacking ...............................................11

10(9′). Slender process extending from middle of apical lobe of genitalia ( Fig. 4e ) ...... ............ S. northlandensis new species

10′. Broad, triangular process extending from middle of apical lobe of genitalia ( Fig. 4h ).......... S. newtoni new species

11(9′). Slender process extending from base of apical lobe of genitalia; minor lobes of left paramere folded ( Fig. 4j )................ ....................... S. dugdalei new species

11′. Apical lobe of genitalia lacking process; minor lobes of left paramere weakly bent ( Fig. 4i ).... S. thorpei new species

12(8′). Slender and short process extending from apical lobe of genitalia; left paramere lacking minor lobe ( Fig. 4d )........ ......... S. coromandelensis new species

12′. Apical lobe of genitalia lacking process; left paramere divided, forming two lobes ...........................................13

13(12′). Apical lobe of genitalia deeply divided into two lobes ( Fig. 4g ) ........................ ....................... S. kuscheli new species

13′. Apical lobe of genitalia not divided.....................................................14

14(13′). Apical lobe of genitalia faucet-shaped; minor lobe of left paramere crescentshaped ( Fig. 4f )..................................... ............... S. taranakiensis new species

14′. Apical lobe of genitalia triangular; minor lobe of left paramere divided but not crescent-shaped....................15

15(14′). Minor lobes of left paramere divided and U-shaped; right paramere rectangular ( Fig. 4m ) .......................................... ................ S. waikatoensis new species

15′. Minor lobes of left paramere divided and V-shaped; right paramere L-shaped ( Fig. 4o )............... S. weiri new species

Figures

Fig. 4. Sagola species in the hirtalis species-group. Aedeagi, dorsal view: a) S. hirtalis, b) S. terricola, c) S. convexa, d) S. coromandelensis, e) S. northlandensis, f) S. taranakiensis, g) S. kuscheli, h) S. newtoni, i) S. thorpei, j) S. dugdalei, k) S. thayerae, l) S. hunuaensis, m) S. waikatoensis, n) S. gisbornensis, o) S. weiri, p) S. brookesi. S. hirtalis: q) Antenna, r) Head, dorsal view, s) Head, ventral view (arrow indicates transverse depression). S. terricola: t) Meso- and metaventrite, u) Male fore femur (arrow indicates semicircular depression). Scale bars = 0.1 mm.

Fig. 5. Known collection localities of New Zealand Sagola species. S. hirtalis: black circles; S. terricola: triangles; S. convexa: black squares; S. coromandelensis: star; S. northlandensis: white circle; S. taranakiensis: white square.

Fig. 6. Known collection localities of New Zealand Sagola species. S. kuscheli: black circle; S. newtoni: triangles; S. thorpei: black squares; S. dugdalei: star; S. thayerae: white circle; S. hunuaensis: white square.

Fig. 7. Known collection localities of New Zealand Sagola species. S. waikatoensis: black circle; S. gisbornensis: triangles; S. weiri: black square; S. brookesi: star.

Fig. 12. Sagola species in the bifida species-group. Aedeagi, dorsal views: a) S. bifida, b) S. latistriata, c) S. crawi, d) S. earlyi, e) S. embersoni, f) S. mayae, g) S. ranatungae, h) S. baylessae, i) S. tekoauensis, j) S. ferroi, k) S. fairburni. S. latistriata, male: l) Dorsal surface of head, m) Ventral surface of head (arrow indicates large depression), n) Mid-femur, lateral view (arrow indicates shallow depression), o) Mid-tibia, lateral view (arrow indicates bend). Scale bars = 0.1 mm.