Sagola newtoni Park and Carlton

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 15-16

Sagola newtoni Park and Carlton , new species

( Figs. 3h , 4h , 6 )

Type Material. Holotype. NEW ZEALAND: Northland: 1♂ ( NZAC), aedeagus dissected and mounted in balsam on a clear plastic card, “ NEW ZEALAND: ND Waipoua State Forest , 0.8km nw Wairau Summit 350m, 27. xi. 1984 hdwd.-podocarp forest A.Newton / M.Thayer 689 ”, “berl., leaf & log litter, forest floor”, “ HOLOTYPE Sagola newtoni Park and Carlton 2013 ” . Paratype (1 male). NEW ZEALAND: Auckland : 1♂ ( DSC), Mt. Auckland, Atuanui Scenic Reserve , 250m, 2.5km NE Glorit, 2 II 2010, D.S. Chandler, sift Nikau palm litter .

Etymology. This species is named for one of the collectors of the holotype and world renowned beetle specialist, Alfred F. Newton.

Diagnosis. This species can be distinguished from the other species of the hirtalis speciesgroup by the following combination of characters: antennomere 1 elongate, approximately 2.5 times longer than wide; posterior frontal fovea elongate; eye small, one-third length of temple; hind wings well-developed; shape of genitalia unique to species.

Description. Length 2.4–2.7 mm. Body brown, elytra, legs, and maxillary palpi yellowish brown ( Fig. 3h ). Head: As long as wide, widest across eyes ( Fig. 3h ). Antennomere 1 approximately 2.5 times longer than wide, 2–3 subquadrate, 4 longer than wide, 5–7 subquadrate, 8–10 transverse. Frontal sulcus deep and reaching hind-point of eye from end of frontal rostral lobes. Anterior frontal fovea round, posterior frontal fovea elongate. Eye small and prominent, one-third length of temple. Thorax: Prosternum as long as wide, widest at one-third length. Elytra rectangular ( Fig. 3h ). Hind wings well-developed. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe longer than parameres with semicircular apical lobe ( Fig. 4h ). Phallobase of median lobe asymmetrical and triangular ( Fig. 4h ). Left paramere longer than right ( Fig 4h ).

Distribution. Auckland, Northland ( Fig. 6 : triangles).

Habitat. Specimens were collected by sifting nikau palm, hardwood, and podocarp forest litter.

Figures

Fig. 3. Habitus of Sagola species in the hirtalis species-group, dorsal views. a) Sagola hirtalis, b) S. terricola, c) S. convexa, d) S. coromandelensis, e) S. northlandensis, f) S. taranakiensis, g) S. kuscheli, h) S. newtoni, i) S. thorpei, j) S. dugdalei, k) S. thayerae, l) S. hunuaensis, m) S. waikatoensis, n) S. gisbornensis, o) S. weiri, p) S. brookesi. Scale bars = 1 mm.

Fig. 4. Sagola species in the hirtalis species-group. Aedeagi, dorsal view: a) S. hirtalis, b) S. terricola, c) S. convexa, d) S. coromandelensis, e) S. northlandensis, f) S. taranakiensis, g) S. kuscheli, h) S. newtoni, i) S. thorpei, j) S. dugdalei, k) S. thayerae, l) S. hunuaensis, m) S. waikatoensis, n) S. gisbornensis, o) S. weiri, p) S. brookesi. S. hirtalis: q) Antenna, r) Head, dorsal view, s) Head, ventral view (arrow indicates transverse depression). S. terricola: t) Meso- and metaventrite, u) Male fore femur (arrow indicates semicircular depression). Scale bars = 0.1 mm.

Fig. 6. Known collection localities of New Zealand Sagola species. S. kuscheli: black circle; S. newtoni: triangles; S. thorpei: black squares; S. dugdalei: star; S. thayerae: white circle; S. hunuaensis: white square.