Sagola valida Broun, 1921, 2013

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 45-47

Sagola valida Broun, 1921

( Figs. 23–24 )

Sagola valida Broun, 1921a: 490 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 244.

Type Material Examined. L e c t o t y p e o f Sagola valida : NEW ZEALAND: Nelson: 1♂ ( BMNH) , glued on rectangular card, “3998. ♂ ” [white label, printed]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “mt. owen. 27.12.1914 ” [white label, handwritten]; “ Sagola valida . ♂.” [white label, handwritten]; “ LECTOTYPE Sagola valida Broun Desig. Park and Carlton 2013 ” [red label, printed]. Paralectotypes (2 males) . NEW ZEALAND: Nelson: 2♂♂ ( BMNH) , Woodhen near Glenhope, T. Hall. The lectotype designation is required because Broun did not explicitly designate a type specimen, and he mentioned that three specimens were examined ( Broun 1921a: 490). This designation will fix the identity of the species and facilitate its recognition by future workers .

Additional Material (n = 59: 28 males; 31 females). NEW ZEALAND: Buller : 2♂♂ 2♀♀, Nelson Lakes NP, Mt. Robert Speargrass Tr. , 875m, 41″49.469′S, 172″48.311′E, 30 XI–17 XII 2005, Nothofagus forest, FMHD#2005-059 , FIT, A. Newton, M. Thayer, ANMT site 1161 ; 1♂, Nelson Lakes NP, Mt. Robert, Speargrass Tr. , 875m, 41″49.469′S, 172″48.311′E, 17 XII 2005, Nothofagus forest, FMHD#2005-110 , berl., leaf & log litter, A. Solodovnikov, D. Clarke, ANMT site 1161 ; 2♂♂ 1♀, Lewis Pass N. R., 11.9km ese Springs Junction , 540m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 715, berl., leaf & log litter, forest floor ; 2♀♀, Lewis Pass N. R., 11.9km ese Springs Junction , 540m, 17 XII 1984 – 21 I 1985, Nothofagus forest , A. Newton, M. Thayer 715, window trap; 1♂ 1♀, Lewis Pass NR, 0.6km s Lewis Pass, 870m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 714, FIT &window trap; 1♂ 1♀, Lewis Pass NR, 13.2km s Lewis Pass, 650m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 713, FIT &window trap; 1♂, Lewis Pass NR, 0.6km s Lewis Pass, 870m, 17 XII 1984 – 21 I 1985, Nothofagus forest, A. Newton, M. Thayer 714, FIT &window trap; 1♂ 1♀, Mt. Robert, carpark, 9 IV 2004, R. Leschen, H. Haman, T. Buckley, litter RL864, 41.49′S, 172.48′E; 1♂, Speargrass Tr., Mt. Robert, 30 XI 2005, J. Nunn; 1♂, Lewis Pass, Asa Pass, 914m, 25 IV 1996, G.W. Ramsay, moss 66/141; 1♂, Rahu Saddle, 670m, 16 X 1984, J.W. Early, moss in Nothofagus forest, LCNZ 84/7; 1♀, Nelson Lakes NP, Lake Rotoiti, St. Arnaud tr., 670m, 14 XII 1984 – 6 I 1985, Nothofagus forest, A. Newton, M. Thayer 706, FIT &window trap; 1♀, Nelson Lakes NP, n slope Mt. Robert, Pinchgut tr., 1290m, 18–26 XII 1984, Noth. Sol. elfin forest, A. Newton, M. Thayer 716, FIT &window trap; 2♀♀, Barrytown Croesus Tr., 10 XI 2005, R. Leschen, S. Nomura, litter RL1022, 42°14′S, 171°19′E; 1♀, Inland Pack tr., 1km n of Punakaiki R, 29 XII 2010, J. Nunn, sifted ground litter; Fiordland: 1♂, Hollyford tr., #110, 44′42S 168′08E, Nothofagus coastal forest leaf litter berlese, 22 I 1998, C. Carlton, R. Leschen; Marlborough: 1♂, Fell Pk, Hut Area, 1300m, Richmond Ra., 13 III 1969, A.C. Eyles, litter 69/109; North Canterbury: 1♂, Arthurs Pass, Mt. Aicken, 950m, 9 II 1984, J.C. Watt, sifted litter 84/24; 1♀, 2.9km N Arthur’ s Pass, Bealey Valley tr., 840–950m, 42′55S 171′33E, #040, under bark, 11 I 1998, C. Carlton, R. Leschen; Nelson: 2♂♂ 2♀♀ (1♂, slide-mounted), Kahurangi NP, Arthur Range, above Flora Saddle, 1000m, 41″11.351′S, 172″44.456′E, 28 XI–19 XII 2005, Nothofagus forest, FMHD #2005-044, FIT, A. Newton, M. Thayer, ANMT site 1156; 1♂ 1♀, Kahurangi NP, Cobb Ridge, above Cobb Reservoir, 1050m, 41″06.351′S, 172″41.658′E, 29 XI–18 XII 2005, Nothofagus cliffortioides forest, FMHD #2005- 051, FIT, A. Newton, M. Thayer, S. Solodovnikov, ANMT site 1159; 1♀, Kahurangi NP, Cobb Ridge, above Cobb Reservoir, 1050m, 41″06.351′S, 172″41.658′E, 18 XII 2005, Nothofagus menziesii & N. solandri cliffortioides forest, FMHD #2005-055, berl., leaf & log litter, M. Thayer, ANMT site 1159; 1♀, Kahurangi NP, Cobb Ridge, above Cobb Reservoir, 1050m, 41″06.351′S, 172″41.658′E, 18 XII 2005, Nothofagus menziesii & N. solandri cliffortioides forest, under bark small Nothofagus logs, M. Thayer, ANMT site 1159; 1♂ 1♀, Kahurangi NP, Cobb Dam Rd., Asbestos Tr., 450m, 41″06.333′S, 172″43,174′E, 29 XI–18 XII 2005, mixed broadleaf-podocarp forest, FMHD #2005-111, berl., leaf & log litter, A. Solodovnikov, D. Clarke, et al., ANMT site 1160; 2♀♀, 0.6km e Gowanbridge, 330m, 18 XII 1984 – 7 I 1985, Nothofagus forest, A. Newton, M. Thayer 717 FIT &window trap, carrion (squid) trap; 1♂ 1♀, Dun Mt., 31 VI 1966, A.K. Walker, litter 66/274; 1♂, Dun Mt., 762m, 13 I 1961, G. Kuschel, litter 61/1; 1♂, Karamea Bluff, 1420ft, J. Nunn; 1♂, Denniston Saddle, 10 II 1999, R. Leschen, R. Hoare, Berl., Nothofagus forest RL288, 41°44′S 171°48′E; 1♂, Arthur Range, w side Flora Saddle, 950m, 1 I 1985, Nothofagus mossy forest, A. Newton, M. Thayer 725, wet leaves & flood debris forest stream; 1♂, Kahurangi NP, Mt. Arthur tr., below Mt. Arthur Hut, 1200m, 28 XI 2005, Nothofagus forest, FMHD #2005-049, berl., leaf litter, A. Solodovnikov, D. Clarke, ANMT site 1157; 1♂, 14 km se Reefton, 250m, 29 V 1982, FMHD #82-606, beech forest litter, S. Peck; 1♀, 30km nw Motueka, Tasman NP, 22 V 1982, FMHD #82-596, beech forest stump litter, S. Peck; 1♂, Dun Mt., 2000′ 14 II 1942, E.S. Gourlay; 1♀, Punakaiki, Pororari Tr., 27 I 1996, J. Nunn, in forest litter; 1♀, Kohaihai River, Karamea, West Coast, 30 V 1963, G. Kuschel, litter; 1♀, Oparara Basin, Box Canyon Cave, 200m, 27 II–1 III 2007, J.W. Early, R.F. Gilbert, Nothofagus podocarp forest, yellow pan trap; Westland: 1♀, Jackson Bay tr., 16 I 2005, R. Leschen, T. Buckley, podocarp, RL906, 43′58′S, 168.37′E; 1♂, 3.2km ne Haast, 14m, Haast River Walk, 43′52S 169′03E, #072, Nothofagus -podocarp forest leaf litter berl., 17 I 1998, C. Carlton, R. Leschen; 1♂, Franz Joseph, 24 II 1989, J. Nunn; 1♀, Hokitika R. Gorge, S Kowhitirangi, 100m, 17 III 1980, podocarp-broadleaf, A. Newton, M. Thayer, berl. leaf & log litter, forest floor; 1♀, 1.8km n Punakaiki, 50m, 19 XII 1984 – 20 I 1985, hdwd.- podo-nikau forest, A. Newton, M. Thayer 719, berl., leaf & log litter, forest floor.

Diagnosis. This species can be distinguished from other Sagola species by the gular region of the male head with small round depression; both sexes with eyes distinctly prominent due to emarginate gena.

Redescription. Length 3.0– 3.5 mm. Body brown, antennae, elytra, legs, and maxillary palpi pale ( Fig. 23a ). Head: Bluntly triangular, widest across eyes ( Fig. 23c ). Male antennomeres 2–3 subquadrate, 4 longer than wide, 5–10 subquadrate. Female antennomeres 2–6 longer than wide, 7–10 subquadrate. Frontal sulcus deep, reaching behind eye from end of frontal rostrum ( Fig. 23c ). Anterior frontal fovea absent ( Fig. 23c ), posterior frontal fovea elongate ( Fig. 23c ). Eye large and prominent, one-half length of temple ( Fig. 23c ). Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular in male ( Fig. 23a ), subquadrate in female. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle in male, patches absent in female. Aedeagus: Median lobe deeply divided, minor lobe longer and narrower ( Fig. 23b ). Phallobase of median lobe symmetrical and rounded ( Fig. 23b ). Parameres symmetrical, ca. 8 setae apically ( Fig. 23b ).

Type Locality. Mount Owen and Woodhen Bend, near Glenhope, Nelson, New Zealand .

Distribution. Buller, Fiordland, Marlborough, North Canterbury, Nelson, Westland ( Fig. 24 : black circles).

Habitat. Most specimens of this species were collected using window and flight intercept traps or by sifting leaf litter or rotten wood in Nothofagus forests. Some specimens were collected from carrion bait or yellow pan traps.

Figures

Fig. 23. Sagola valida (valida species-group). a) Habitus, dorsal view, b) Aedeagus, dorsal view, c) Dorsal surface of male head, d) Ventral surface of male head. Scale bar a = 1 mm, b–d = 0.1 mm.

Fig. 24. Known collection localities of Sagola valida (black circles).