Sagola anisarthra Broun, 1893, 1053

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 78-80

Sagola anisarthra Broun, 1893

( Figs. 45a, 45f, 45 l-m, 46)

Sagola anisarthra Broun, 1893b: 1053 . Raffray 1904: 498, 1911: 6, 1924: 233; Hudson 1923: 365, 1934: 183; Newton and Thayer 2005b; Nomura and Leschen 2006: 241.

Sagola suturalis Broun, 1914b: 158 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 244. New synonymy.

Sagola dickensis Broun, 1917: 377 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242. New synonymy.

Type Material. Holotype. NEW ZEALAND: Dunedin : 1♂ ( BMNH), glued on rectangular card, “Sharp Coll. 1905-313.” [white label, printed]; “ Sagola anisarthra fide Sandager Moeraki” [white label, handwritten] . Syntypes of Sagola dickensis : NEW ZEALAND: Otago Lakes : 1♂ ( BMNH), glued on rectangular card, “Type” [red label, printed], “3827. ♂ ” [white label, handwritten], “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed], “Mt. Dick. 10.3.1914 ” [white label, handwritten], “ Sagola . ♂ dickensis.” [white label, handwritten]; 1♂ ( BMNH), glued on rectangular card, “3827. ♂ ” [white label, handwritten]; “ New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Mt. Dick. 10.3.1914 ” [white label, handwritten]; “ Sagola . ♂ dickensis.” [white label, handwritten]; 1♂ ( NZAC), glued on rectangular card, “Coll. T.Hall. 10-3-1914.” [white label, handwritten], “Mt. Dick, W. of L. Wakatipu, Southland.” [white label, handwritten], “3827.” [white label, handwritten], “ Paratype.” [white label, handwritten], “ Sagola dickensis Broun. ” [white label, handwritten], “ T.Broun Collection” [white label, handwritten], “A.E.Brookes Collection” [white label, handwritten], “N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand” [yellow label, printed]; 1♂ ( NZAC), glued on rectangular card and aedeagus dissected and mounted in balsam on a clear plastic card, “ Sagola dickensis Broun. ” [white label, handwritten], “3827.” [white label, handwritten], “L. Wakatipu Region, Southland.” [white label, handwritten], “Coll. T.Hall, march, 1914.” [white label, handwritten], “A.E.Brookes Collection” [white label, handwritten], “T.Broun Collection” [white label, handwritten], “N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand” [yellow label, printed] ; 1♂ ( NZAC), glued on rectangular card, “Coll. T.Hall, march, 1914.” [white label, handwritten], “L. Wakatipu Region, Southland.” [white label, handwritten], “3827.” [white label, handwritten], “ Sagola dickensis Broun. ” [white label, handwritten], “T.Broun Collection” [white label, handwritten], “A.E.Brookes Collection” [white label, handwritten], “N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand ” [yellow label, printed] ; 2♂♂ ( NZAC), glued on rectangular card, “Coll. T.Hall, march, 1914.” [white label, handwritten], “L. Wakatipu Region, Southland.” [white label, handwritten], “3827.” [white label, handwritten], “ Sagola dickensis Broun. ” [white label, handwritten], “T.Broun Collection” [white label, handwritten], “A.E.Brookes Collection” [white label, handwritten], “N.Z. Arthropod Collection, NZAC Private Bag 92170 AUCKLAND New Zealand ” [yellow label, printed]. *Original description examined eight specimens, but we could not find one of them. Syntypes of Sagola suturalis : NEW ZEALAND: Mid Canterbury : 1♂ ( BMNH), glued on rectangular card, “type” [red label, printed], “3522. ♂ ” [white label, handwritten], “ Rakaia. 5.6.1912 ” [white label, handwritten], “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed], “ Sagola . ♂ suturalis.” [white label, handwritten] ; ♂, glued on rectangular card, “3522. ♂ ” [white label, handwritten], “ Rakaia Gorge. 5.6.1912 ” [white label, handwritten], “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed], “ Sagola . ♂. suturalis.” [white label, handwritten] .

Additional Material (n = 37: 30 males; 7 females). N E W Z E A L A N D: D u n e d i n: 10♂♂ (1♂, slide-mounted), Boulder Beach, Otago Penin. , 19 XI 2004, washed up live with incoming tide, J. Nunn ; 1♀, Mt. Watkin nr Waikouaiti , XII 2007, pit trap, Kelvin Lloyd ; 1♂, Boulder Beach , Otago Penin. , 7 XII 2002, washed up live with incoming tide, J. Nunn ; 1♂, Boulder Beach , Otago Penin. , 14 I 2005, washed up live with incoming tide, J. Nunn ; 2♂♂, Mt. Cargill, Grahams Bush , 260m, 45″48.299′S 170″34.905′E, 14 XII 2005, mixed broadleaf forest on steep slope w/emergent podocarp, FMHD#2005-107 , litter, A. Newton, M. Thayer, ANMT site 1164; Fiordland: 4♂♂ 1♀, S of Lake Hauroko , 12 XI 1966, F. Alack, litter 66/420; Mid Canterbury : 4♂♂ 2♀♀, Goat Hill, nr. Metheven, T. Hall , 20 X 1913 ; 1♂, Mt Hutt , 11 V 2003, A.C. Eyles, mountain beech litter; Southland : 1♀, Clifden, Limestone Bluffs , 11 II 1968, J.I. Townsend, moss ; 5♂♂ 1♀, MW relay stn, Blue Mountains , 19 III 2006, J. Nunn ; 1♂, Lookout Point Bluff , 7 VII 2001, amongst tussock grasses, J. Nunn ; 1♀, Pounawea Catlins , 25 X 2003, litter at beach streamline, J. Nunn .

Diagnosis. This species can be distinguished from the other species of the anisarthra speciesgroup by the following combination of characters: larger body, length 2.4–2.8 mm; male antennomeres 4–6 enlarged and 4–11 bearing tubercles; frontal sulcus deep reaching midpoint of eye; anterior frontal fovea round and partially covered by frontal rostrum, posterior frontal fovea oval; eye large and prominent, approximately twothirds length of temple; shape of genitalia unique to species.

Redescription. Length 2.4–2.8 mm. Body reddish brown, antennae, legs, maxillary palpi, and elytra paler ( Fig. 45a ). Head: Bluntly triangular, widest across eyes ( Fig. 45l ). Ventral surface of head weakly convex. Male antennomere 1 approximately 2 times longer than wide, 2–10 subquadrate, 4–6 enlarged, 7–10 bearing tubercles. Female antennomeres 2–3 subquadrate, 4–5 longer than wide, 6–10 subquadrate, 4–11 bearing tubercles. Frontal sulcus deep, reaching midpoint of eye. Anterior frontal fovea round and partially covered by frontal rostrum ( Fig. 45l ), posterior frontal fovea oval ( Fig. 45l ). Eye large and prominent, approximately one-half length of temple ( Fig. 45l ). Thorax: Prosternum as long as wide, widest at midpoint. Elytra rectangular ( Fig. 45a ). Hind wings well-developed. Meso- and metathorax trapezoidal, longer than wide. Abdomen: Tergite IV with pair of transverse patches of microtrichia reaching middle. Aedeagus: Median lobe as long and wide as parameres ( Fig. 45f ). Phallobase of median lobe symmetrical and rounded ( Fig. 45f ). Parameres symmetrical with setae apically ( Fig. 45f ).

Type Locality. Moeraki , Dunedin, New Zealand .

Distribution. Dunedin, Fiordland, Mid Canterbury, Otago Lakes, Southland ( Fig. 46 : black circles).

Habitat. Most specimens of this species were collected by sifting mossy leaf or log litter in forests or the intertidal zone of beach areas.

Comments. Specimens of S. anisarthra can be distinguished from other species by the enlarged male antennomeres 4–6 and dorsal shape of head. The type specimens of S. dickensis and S. suturalis share these diagnostic characters, and additionally those species have been collected at or near the type localities. For these reasons, we have placed S. dickensis and S. suturalis in synonymy with S. anisarthra .

Figures

Fig. 45. Sagola species in the anisarthra species-group. Habitus, dorsal views: a) S. anisarthra, b) S. planipennis, c) S. townsendi, d) S. wairarapaensis, e) S. clarkei. Aedeagi, dorsal views: f) S. anisarthra, g) S. planipennis, h) S. planipennis, lateral view, i) S. townsendi, j) S. wairarapaensis, k) S. clarkei. S. anisarthra, male head: l) Dorsal surface, m) Ventral surface. Scale bars a–e = 1 mm, f–m = 0.1 mm.

Fig. 46. Known collection localities of New Zealand Sagola species. S. anisarthra: black circles; S. planipennis: triangles; S. townsendi: black square; S. wairarapaensis: star; S. clarkei: white circles.