Sagola hectorii Broun, 1917

Park, J. - S. & Carlton, C. E., 2014, A revision of the New Zealand species of the genus Sagola Sharp (Coleoptera: Staphylinidae: Pselaphinae: Faronitae), The Coleopterists Bulletin (mo 13) 68, pp. 1-156 : 128

Sagola hectorii Broun, 1917

( Figs. 68b , 69b , 70b , 71 )

Sagola hectorii Broun, 1917: 378 . Hudson 1923: 365, 1934: 184; Newton and Thayer 2005b; Nomura and Leschen 2006: 242.

Sagola distorta Broun, 1921b: 602 . Hudson 1923: 366, 1934: 184; Newton and Thayer 2005b. Nomura and Leschen 2006: 242. New synonymy.

Type Material. Holotype. NEW ZEALAND: Nelson: 1♂ ( BMNH), glued on rectangular card, “Type” [red label, printed]; “3829.” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed]; “Belgrove. 7.11.1914.” [white label, written]; “ Sagola . ♂ hectori” [white label, handwritten] . Holotype of Sagola distorta . NEW ZEALAND: Wairarapa: 1♂ ( BMNH), glued on rectangular card, “4166. ♂ ” [white label, handwritten]; “New Zealand Broun Coll. Brit. Mus. 1922–482.” [white label, printed] ; “Featherston. 6.10.1916.” [white label, written]; “ Sagola distorta . ♂ ” [white label, handwritten].

Additional Material (n = 8: 3 males; 5 females). NEW ZEALAND: Nelson : 1♂ 2♀♀, Upper Maitai , 7 IV 1963, E.S. Gourlay ; 1♂, Upper Maitai , 7 III 1949, E.S. Gourlay ; 3♀♀, Upper Maitai , 3 V 1950, E.S. Gourlay ; 1♂, Dovedale , 11 X 1963, J.I. Townsend, litter .

Diagnosis. This species can be distinguished from the other species of the insignis species-group by the following combination of characters: body length 2.4–2.8 mm; temporal lobe of male head distinctly projecting, exceeding eye, with weakly concave ventral surface; male temporal lobe with L-shaped depression anteriorly; ventral surface of male head with transverse process behind mouthparts with depression medially and setae along edge; elytra subquadrate; shape of antennomeres and genitalia unique to species.

Redescription. Length 2.4–2.8 mm. Body brown, antennae, legs, maxillary palpi, and elytra paler ( Fig. 68b ). Head: Male head triangular, widest across temples ( Fig. 69b ). Female head bluntly triangular, widest across eyes. Antennomere 1 approximately 1.5 times longer than wide, 2–3 subquadrate, 4–7 longer than wide, 8–10 subquadrate. Frontal sulcus deep, reaching end of eye. Anterior frontal fovea round, partially covered by frontal rostrum, posterior frontal fovea round. Male eye prominent, approximately one-fourth length of temple, female slightly smaller. Temporal lobe of male head distinctly projecting, exceeding eye, with weakly concave ventral surface ( Fig. 69b ). Male temporal lobe with L-shaped depression anteriorly ( Fig. 69b ). Ventral surface of male head with transverse process behind mouthparts with depression medially, with setae along edge ( Fig. 69b ). Thorax: Prosternum as long as wide, widest at one-third length. Elytra subquadrate ( Fig. 68b ). Meso- and metathorax trapezoidal, as long as wide. Abdomen: Tergite IV without patches of microtrichia. Aedeagus: Median lobe of genitalia broad with triangular head ( Fig. 70b ). Phallobase of median lobe symmetrical and rounded ( Fig. 70b ). Parameres symmetrical and triangular, setose from midpoints to apices ( Fig. 70b ).

Type Locality. Belgrove , near Nelson, New Zealand .

Distribution. Nelson ( Fig. 71 : triangles).

Habitat. One specimen was collected by sifting leaf litter.

Comments. Specimens of S. hectorii can be distinguished from those of other species by the shape of the ventral surface of the male head and the antennomeres. The type specimen of S. distorta shares these diagnostic characters. For these reasons, we have placed S. distorta in synonymy with S. hectorii .

Figures

Fig. 68. Habitus of Sagola species in the insignis species-group, dorsal views. a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 1 mm.

Fig. 69. Ventral views of male heads of Sagola species in the insignis species-group. a) Sagola insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Scale bars = 0.1 mm.

Fig. 70. Sagola species in the insignis species-group. Aedeagi, dorsal views: a) S. insignis, b) S. hectorii, c) S. bipunctata, d) S. angulifera, e) S. eminens, f) S. robustula, g) S. pertinax, h) S. monticola, i) S. laticeps, j) S. castanea, k) S. sheldoni, l) S. mercuryensis, m) S. cuvierensis, n) S. buckleyi, o) S. brumfieldi, p) S. otteai, q) S. boudreauxae, r) S. taegyui, s) S. hanae, t) S. gyudongi, u) S. egmontensis, v) S. boonei, w) S. stewartensis, x) S. jeongnamae. Male heads, dorsal surface: y) S. bipunctata, z) S. eminen. Scale bars = 0.1 mm.

Fig. 71. Known collection localities of New Zealand Sagola species. S. insignis: black circles; S. hectorii: triangles; S. bipunctata: black squares; S. angulifera: stars; S. eminens: white circles; S. robustula: white squares.