Calloeneis roxanne Gordon and Hanley, 2020

Gordon, Robert D., F, Guillermo González & Hanley, Guy A., 2020, South American Coccinellidae (Coleoptera), part XX: systematic revision of South American Calloeneis Grote (Cryptognathini), Insecta Mundi 2020 (766), pp. 1-25 : 11

11. Calloeneis roxanne Gordon and Hanley , new species

Description. Male holotype. Length 1.6 mm, width 1.3 mm; dorsal surface shiny, head distinctly alutaceous. Color yellow; elytron with narrow, oval dark brown macula on sutural margin in basal 1/2, additional black macula present on lateral margin in basal 1/2 ( Fig. 43 ); meso,- metasternum black; abdomen with basal ventrite dark brown. Head punctures separated by a diameter or less; pronotal punctures small, separated by up to twice a diameter; elytral punctures larger than on pronotum, separated by up to three times a diameter; prosternal punctures small, separated by about a diameter; mesosternal punctures larger than on prosternum, separated by less than a diameter, metasternal punctures large, separated by a diameter or less medially, absent in lateral 1/3; abdominal ventrites 1, 2 with fine punctures separated by up to 3 times diameter medially, ventrites 2–4 finely, densely punctured medially; ventrite 5 finely, densely punctured. Head with frons parallel sided, 1.2 times as wide as eye, medially depressed. clypeal apex truncate ( Fig. 44 ); eye canthus short, about half width of eye. Prosternum with lateral carina on each side curved, extended from apex of intercoxal process to lateral margin of prosternum. Epipleuron not descending externally. Postcoxal line on basal abdominal ventrite curved, extended 3/4 distance to rear margin of ventrite. Genitalia with penis guide shorter than paramere, sides parallel in basal 3/4, rounded to acute apex; paramere slender, curved, apex abruptly rounded ( Fig. 45, 46 ); penis long, slender, apex shortly attenuate, basal capsule lost ( Fig. 47 ).

Female. Similar to male except genitalia with ramus bulbous, cornu strongly narrowed from base to slender apex ( Fig. 48 ).

Variation. Length 1.5 to 2.0 mm, width 1.3 to 1.7 mm. Elytral color pattern varies from that described above to having lateral macula absent or having both maculae absent.

Type material. Holotype male: Chapada, Brazil, Acc No. 2966, Aug. ( CMNH) . Paratypes 29, same data as holotype except months April, June, September, October, November. ( CMNH).

Remarks. This species has a recognizable dorsal color pattern, although the pattern is somewhat variable. Otherwise male genitalia must be examined for identification.

Figures

Figures 38–53. Calloeneis spp.38–42) Calloeneis krista. 38) Habitus.39) Frons.40) Penis guide ventral.41) Penis guide lateral. 42) Penis. 43–48) Calloeneis roxanne. 43) Habitus. 44) Frons. 45) Penis guide ventral. 46) Penis guide lateral. 47) Penis. 48) Spermathecal capsule. 49–53) Calloeneis angelica. 49) Habitus. 50) Frons. 51) Penis guide ventral. 52) Penis guide lateral. 53) Penis.