Calloeneis Grote, 1873

Gordon, Robert D., F, Guillermo González & Hanley, Guy A., 2020, South American Coccinellidae (Coleoptera), part XX: systematic revision of South American Calloeneis Grote (Cryptognathini), Insecta Mundi 2020 (766), pp. 1-25 : 2

Calloeneis Grote

Calloeneis Grote 1873: 143 ; Belicek 1976. Replacement name for Oeneis Mulsant.

Oeneis Mulsant 1850: 497 , 500. Not Hübner 1819; Gordon 1971: 184.

Cryptognatha Mulsant : Crotch 1874: 206 (in part); Blackwelder 1945: 449 (in part); Gordon 1971: 184. Delphastopsis Casey 1924: 170 ; Gordon 1971: 184.

Dargo Chapin 1955: 87 . Unnecessary replacement name for Oeneis Mulsant ; Gordon 1971: 184.

Type species. Oeneis nigrans Mulsant , by subsequent designation of Korschefsky (1931).

Description. Cryptognathini with anterior margin of clypeus margin weakly arcuate or truncate, anterior angle slightly upturned (Fig. 95); clypeus slightly expanded onto eye; antenna with eight articles, club apparently with three articles; maxillary palpus with apical article widened slightly apically. Prosternum approximately as long as median 1/3 of basal abdominal ventrite, wide anterior to coxae, completely concealing mouthparts in repose (Fig. 97); prosternal carina long, extended from apex of prosternal process to lateral margin of prosternum. Epipleuron weakly descending externally or flat (Fig. 96). Proleg with tibia moderately expanded for reception of tarsus, slightly narrower than femur, femur notched for reception of tibia. Meso- and metatibia less strongly expanded, grooved on outer margin for reception of tarsi, femur narrow, notched for reception of tibia. Abdomen with postcoxal line incomplete. Apex of male genitalia arcuate, unmodified. Female genital plate with stylus and stiff setae; spermatheca variable ( Fig. 5 , 16 , 37 ).

Remarks. Calloeneis is clearly defined by key characters. It is most similar to Cryptognatha but differs from that genus by a mostly unmodified clypeus, slightly descending or nearly flat epipleuron, and slightly less strongly expanded prosternum. Korschefsky (1931) designated Oeneis nigrans as the type species of Oeneis Mulsant. However , of the two species included in that genus by Korschefsky (1931), O. obscura Mulsant and O. nigrans , type depositories could not be located so neither is included in this study. A type specimen of Cryptognatha nitidella Brèthes in the BMNH was examined and determined to not belong to Cryptognathini . It is a species of Coccinellidae , but generic provenance was not recognized.

Figures

Figures 1–13. Calloeneis spp. 1–6) Calloeneis lynne. 1) Habitus. 2) Frons. 3) Penis guide ventral. 4) Penis guide lateral. 5) Penis. 6) Spermathecal capsule. 7–11) Calloeneis sheri. 7) Habitus. 8) Frons. 9) Penis guide ventral. 10) Penis guide lateral. 11) Penis. 12–13) Calloeneis marianne. 12) Habitus. 13) Frons.

Figures 14–25. Calloeneis spp. 14–16) Calloeneis kara. 14) Habitus.15) Frons.16) Spermathecal capsule.17–20) Calloeneis jacquelin. 17) Habitus. 18) Frons. 19) Spermathecal capsule. 20) Habitus variation. 21–25) Calloeneis blanca. 21) Habitus. 22) Frons. 23) Penis guide ventral. 24) Penis guide lateral. 25) Penis.

Figures 26–37. Calloeneis spp. 26–29) Calloeneis myra. 26) Habitus.27) Frons.28) Penis guide ventral. 29) Penis guide lateral. 30–31) Calloeneis leticia. 30) Habitus. 31) Frons. 32–37) Calloeneis signata. 32) Habitus. 33) Frons. 34) Penis guide ventral. 35) Penis guide lateral. 36) Penis. 37) Spermathecal capsule.