Calloeneis bethany Gordon and Hanley, 2020

Gordon, Robert D., F, Guillermo González & Hanley, Guy A., 2020, South American Coccinellidae (Coleoptera), part XX: systematic revision of South American Calloeneis Grote (Cryptognathini), Insecta Mundi 2020 (766), pp. 1-25 : 16-17

21. Calloeneis bethany Gordon and Hanley , new species

Description. Male holotype. Length 1.6 mm, width 1.5 mm; dorsal surface shiny except head alutaceous. Color black; head yellow; pronotum yellow with small, dark brown basomedian macula narrowly extended to apical margin ( Fig. 91 ); venter yellow except prosternum, mesosternum black, epipleuron brownish yellow. Head with punctures not visible; pronotal punctures small separated by less than to twice a diameter; elytral punctures larger than on pronotum, separated by less than to twice a diameter; prosternal punctures fine, barely visible; mesosternal punctures coarse, separated by less than a diameter, metasternal punctures large, separated by a diameter or less, absent in lateral 1/3; abdominal ventrite 1 impunctate medially except some large punctures on anterior 1/4, ventrites 2–4 with fine punctures not clearly visible; ventrite 5 finely, densely punctured. Head with frons parallel sided, 1.2 times as wide as eye, clypeal apex slightly arcuate ( Fig. 92 ); eye canthus short, about half width of eye. Prosternum with lateral carina on each side curved, extended from apex of intercoxal process to lateral margin of prosternum. Epipleuron not descending externally. Postcoxal line on basal abdominal ventrite curved, extended to rear margin of ventrite. Genitalia with penis guide longer than paramere, basally narrow, strongly widened from base to median 1/2, then abruptly widened, side rounded, tapered to acute apex in apical 1/4 ( Fig. 93, 94 ); penis lost.

Female. Similar to male except for the black head and reddish-brown clypeus.

Variation. Length 1.4 to 1.6 mm, width 1.3 to 1.5 mm. Male pronotal macula may be slightly larger than described above.

Type material. Holotype male: Peru, Satipo , IX-X, 1942, Paprzycki ( USNM) . Paratypes, 4, same data as holotype except dates X, 1942, XI, 1942 ( USNM).

Remarks. Male genitalia are distinctive, but a mostly yellow pronotum and Peruvian type locality will also aid in identification.

Figures

Figures 81–94. Calloeneis spp. 81–85) Calloeneis alexandra. 81) Habitus. 82) Frons. 83) Penis guide ventral. 84) Penis guide lateral. 85) Penis. 86–90) Calloeneis brooke. 86) Habitus. 87) Frons. 88) Penis guide ventral. 89) Penis guide lateral. 90) Penis. 91–94) Calloeneis bethany. 91) Habitus. 92) Frons. 93) Penis guide ventral. 94) Penis guide lateral.