Austropsyche bifurcata (Kimmins)

Wells, Alice & Contents, Arturs Neboiss Table Of, 2018, Australian Diplectroninae reviewed (Insecta: Trichoptera), with description of 21 new species, most referred to a new genus, Zootaxa 4415 (1), pp. 1-44 : 19

Austropsyche bifurcata (Kimmins)

( Figs 49 , 63–65, 71 , 88–89, 96–97)

Diplectrona bifurcata Kimmins, in Mosely & Kimmins 1953 , 344. Austropsyche bifurcata ; Neboiss 1986, 219.

Material examined. Holotype ♂, Mt Kosciusko, 5,000 ft, NSW ( BMNH) ; paratype 1♂, same data as for holotype ( ANIC). ACT: 1♂, Bendora, 14.xii.1960, D. Colless [PT-1015] ; 3♂ 2 larvae, Mt Gingera, 11.i.1967, E.F. Riek (ANIC). New South Wales: 2♂ 4♀, Dead Horse Gap, 5.i.1984, G. Theischinger [TRI-13803] . Victoria: 1♂, Kanuka Creek, Helmers Road, 420 m, East Gippsland , 11.iii.1982, J. Blyth [PT-1008].

Diagnosis. Males of Austropsyche bifurcata belong in the Au. victoriana Group; Au. bifurcata is distinguished from Au. victoriana by the absence in male genitalia, in ventral view, of the ventrolateral plates at the base of the gonopods and differs from Au. kaputar in having the mesal process on each coxopodite subapical, and short, not basal, elongate, and stout.

Description. Length of each forewing: ♂ 9.0–10.0 mm (n = 3), ♀ 11.5–12.8 mm (n = 3); median cell length about 1.3x length discoidal cell.

Male. Genitalia ( Figs 63–65 , 88–89 ): Tergite IX reduced to narrow bridge midventrally, in lateral view partly delineated from tergite X, with small convexity below phallic apparatus midapicolaterally, rounded in ventral view; abdominal tergite X apically with deep V-shaped excision; gonopods each with coxopodite about 1.5–2.0x length of harpago and well-developed irregular-shaped mesal lobe subapically; phallic apparatus with two pairs of sharply pointed, straight endothecal spines subapically.

Female (association tentative, Figs 96, 97 ). Resembling Au. victoriana : Terminal abdomen with sternite VIII divided to form 2 plates, in ventral view broad based, tapered distally to truncate apices, in lateral view, apicolateral angles slightly produced, triangular.

Distribution. Collected at higher altitudes of south-eastern Australia from East Gippsland, Victoria to the Blue Mountains, west of Sydney, New South Wales.

Figures

FIGURES 49–58. Austropsyche Banks 1939 spp., diagnostic features. 49–50, ♂ right fore- and hind wings, dorsal: 49, Au. bifurcata (Kimmins 1953; holotype); 50, Au. ambigua sp. nov. (NSW, Upper Manning River, 20 km NNW Rawdon Vale [PT- 1019]). 51, Au. ambigua sp. nov. [PT-1019], ♂ maxillary palp. 52–56, Au. victoriana Banks 1939, ♂ genitalia: 52, (holotype), left lateral; 53–54, (Vic., Kalymna Falls, Williams Range, Grampians [PT-1009]): 53, ventral, and 54, left lateral; 55–56, (Vic., CementCreek [PT-1100]):55, left lateral,and 56, phallic apparatus, leftlateral and ventral and dorsal, respectively, leftto right. 57–58, ♀ terminal abdominal segments, left lateral: 57, Au. victoriana (Vic, Cumberland Falls, SE Marysville [PT-1046]); 58, Au.ambigua sp. nov. (NSW, Upper Manning River, 20 km NNW Rawdon Vale [PT-1058]). Scale bars = 2 mm.Abbreviations: ac = anal cell; A 1+2+3 = fused anal veins A 1, A 2, A 3;dc = discoidal cell; cox = coxopodite; coxp = mesal process on coxopodite; fI, fII, fIII, fIV, fV = forks I–V; mc = median cell; har = harpago; ph = phallic apparatus; VIII, IX, X = abdominal segments VIII, IX, X.

FIGURES 70–83. Austropsyche Banks 1939 spp., ♂ diagnostic features.70–75, heads, dorsal:70, Au.victoriana Banks 1939; 71, Au.bifurcata (Kimmins 1953); 72, Au. kaputar sp. nov.; 73, Au. acuta sp. nov.; 74, Au. ambigua sp. nov.; 75, Au. morana sp. nov. 76–79, right fore- and hind wings, dorsal: 76, Au.victoriana; 77, Au. kaputar sp. nov.; 78, Au.acuta sp. nov.; 79, Au. morana sp. nov. 80, Au. acuta sp. nov., antennal segments. 81, Au. acuta sp. nov., maxillary palp. 82, Au. victoriana, reticulate-walled internal sacs of abdominal segments VI and VII, ventral. 83, Au. acuta sp. nov., left lateral filament on segment V, ventral. Scale bars = 2 mm. Abbreviations: ac = anal cell; alw = anterolateral wart; amw = anteromesal wart; A 1+2+3 = fused anal veins A 1, A 2, A 3; dc = discoidal cell; mc = median cell; pw = posterior wart.

FIGURES 59–69. Austropsyche Banks 1939 spp., ♂ genitalia diagnostic features. 59–60, Au. acuta sp. nov. (holotype, NSW, Crystal Creek Rainforest Resort, Upper Crystal Creek; ANIC): 59, ventral; 60, left lateral.61–62, Au. ambigua sp. nov. (NSW, Upper Manning River,20 km NNW Rawdon Vale [PT-1019]):61, ventral; 62, leftlateral.63–65, Au.bifurcata (Kimmins1953; ACT, Bendora ([PT-1015]), ♂ genitalia: 63, ventral; 64, apex of phallic apparatus, left lateral; 65, left lateral. 66–67, Au. morana sp. nov. (Qld, trickle on Moran's Falls track, Lamington NP [TRI-54706-1]): 66, ventral; 67, left lateral. 68–69, Au. kaputar sp. nov. (NSW, Mt Kaputar NP, Bullawa Creek [PT-1714]): 68, ventral; 69, left lateral.

FIGURES 84–101. Austropsyche Banks 1939 spp., genitalia diagnostic features. 84–95, ♂ genitalia: 84–87, Au. victoriana Banks 1939:84–85, (typical form): 84, ventral, and 85, left lateral; 86, (Vic., Wilhelmina Falls [TRI-12958]), ventral, showing variation in development of the ventrolateral ‘plates’ at the bases of the gonopods; 87, (Vic., Cumberland Falls, SE Marysville [PT-1018]), ventral, showing variation in development of the ventrolateral ‘plates’ at the bases of the gonopods. 88–89, Au. bifurcata (Kimmins 1953; (ACT, Bendora [1015]): 88, ventral, and 89, left lateral. 90–91, Au.kaputar sp. nov. (holotype): 90, ventral, and 91, left lateral. 92, Au. acuta sp. nov. (NSW, Upper Crystal Creek), ventral. 93, Au. ambigua sp. nov. (Vic. Noorinbee [1097]), ventral.94–95, Au. morana sp. nov. (holotype): 94, ventral; 95, left lateral.96–101, ♀ terminal abdominal segments.96–97, Au.bifurcata (NSW, New South Wales,Dead Horse Gap): 96, ventral, and 97, left lateral; 98, Au.victoriana, ventral. 99, Au. morana sp. nov. (Qld, trickle on Moran's Falls track, Lamington NP), ventral. 100–101, Au. acuta sp. nov. (NSW, Upper Crystal Creek): 100, ventral, and 101, left lateral.