Conexicoxa Lin, 1986

Strelnikova, O. D. & Yan, E. V., 2023, ON SPLITTING OF THE GENUS NOTOCUPES (COLEOPTERA: ARCHOSTEMATA): NEW DATA ON MORPHOLOGY AND TAXONOMY, Far Eastern Entomologist 488, pp. 1-28 : 15-16

Genus Conexicoxa Lin, 1986 , stat. resurr.

Type species: Conexicoxa homora Lin, 1986 .

DIAGNOSIS. Beetles with narrow (body length to width ratio is 2.7–2.9) or wide body (body length to width ratio is 1.7–2.2). Integument with single type of medium-size (0.02–0.04 mm in diameter), or, less common, small (0.01–0.02 mm in diameter) tubercles ( Fig. 29 ). Antennae filiform or weakly serrated ( C. denticollis ). Pronotum trapezoid, pronotal disc bears mesal cordate protuberance. Elytral humerus sloping, not protruding laterally (except С. psilata ), epipleural margin gradually curving along its length, epipleuron wide (except in С. siniestri and С. exigua which have narrow epipleuron), gradually, weakly narrowing or almost not narrowing apically, elytral veins 2 and 3 curving towards sutural margin, not forming straight parallel rows, could merge together before reaching elytral apex. Abdomen not narrowing basally ( Fig. 58 ).

REMARKS. Conexicoxa was previously treated as a junior synonym of Notocupes ( Ponomarenko et al., 2012) , but restored as separated genus in the present study for Notocupes species with trapezoid pronotum and curved epipleural margin.

COMPOSITION. This genus consists of 17 species from Jurassic of Germany, Mongolia, China and Kyrgyzstan, Upper Jurassic-Lower Cretaceous of China, Lower Cretaceous of Spain and Russia, namely С. homora Lin, 1986 , С. brachicephala (Ponomarenko, 1994) , comb. n. [ Notocupes ], С. crassa ( Ponomarenko, 1969) , comb. n. [ Notocupes ], C. denticollis ( Jiang et al., 2020) , comb. n. [ Notocupes , Echinocups ], С. epicharis ( Tan, Ren et Liu, 2005) , comb. n. [ Amblomma , Notocupes ], С. eumeura (Tan et al., 2006), comb. n. [ Amblomma , Notocupes ], С. exigua (Ponomarenko, 1994), comb. n. [ Notocupes ], С. jurassica (Hong, 1983), comb. n. [ Chengdecupes , Notocupes ], С. khetanensis (Ponomarenko, 1993), comb. n. [ Notocupes ], С. kirghizica ( Ponomarenko, 1969) , comb. n. [ Notocupes ], С. longicollis (Ponomarenko, 1994) , comb. n. [ Notocupes ], С. martinclosas (Soriano et Martinez-Delclòs, 2006), comb. n. [ Zygadenia , Notocupes ], С. porrecta (Tan et al., 2006), comb. n. [ Amblomma , Notocupes ], С. psilata ( Tan, Ren et Liu, 2005) , comb. n. [ Amblomma , Notocupes ], С. siniestri (Soriano et Martinez-Delclòs, 2006), comb. n. [ Zygadenia , Notocupes ], С. stabilis ( Tan, Ren et Liu, 2005) , comb. n. [ Amblomma , Notocupes ], С. tripartita (Oppenheim, 1888), comb. n. [ Procarabus , Notocupes ].

Figures

Figs 29–31. Characteristic of cuticular coverings five investigated genera of Archostemata. 29 – Rhabdocupes oxypygus, covered with one type of cuticular tubercles; 30 – Notocupes caudatus, covered with two types of cuticular tubercles; 31 – Brachilatus caducus, covered with three types of cuticular tubercles. Scale bar = 1 mm.

Figs 57–60. Abdomen morphology. 57 – Rhabdocupes rostratus; 58 – Conexicoxa longicollis; 59 – Brachilatus caducus; 60 – Notocupes lapidarius. Scale bar = 1 mm.