Orbellis longirostris Zámbó & Kondorosy, 2023

Zámbó, András, Kovács, Szilvia & Kondorosy, Előd, 2023, Revision of the genus Orbellis Distant, 1913 (Heteroptera: Rhyparochromidae), Zootaxa 5389 (2), pp. 193-212 : 208-209

Orbellis longirostris Zámbó & Kondorosy , sp. nov.

( Figs. 11 , 28 )

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Type material. HOLOTYPE: Madagascar 2011 / Ambohitantely Spec. Res. / S 18°11’51” E 47°17’03” / 1530m; at light, 24–29. xi / M.Trýzna lgt. (♀, Collection Ernst Heiss, Innsbruck, Austria). Type locality. Ambohitantely Spec. Res

Description.

Colour: Head fuscous; scape brown except yellow base; pedicel except fuscous apex, base of basiflagellum and labium yellow; apical one-third of basiflagellum cream coloured. Thorax fuscous with all punctures as well; lateral spots of pronotum confluent with fuscous ground colour and lateral keels of scutellum brown; very narrowly anterior and whole lateral margin of pronotum, three spots on posterior margin of pronotum (narrowly confluent on very margin), spots of hemelytra and lateral margin of corium except brown spots in posterior 1/3 and legs (except fuscous femora and brown apex of tibiae and tarsomeres III) yellow; hemelytra mostly yellow with fuscous spots as follows: between veins R+M and Cu in basal one-fourth, between veins R+M at branching, a rounded one between base of M and Cu continuing on both sides of M to apical margin, inner and lateral angle of apical margin of corium, and some diffuse spots (most characteristic spots on inner part of anterior half of clavus). Membrane amber. Abdomen fuscous.

Structure: Body dull, head, sternum, antenna and legs more or less shiny. Dorsal surface of head slightly coriaceous and with minute deep punctures, ventral part sparsely punctate with minute punctures. Labium surpassing middle of abdominal sternite V. Pronotal collar relatively wide (about 3 times as thick medially as base of pedicel), slightly V-shaped, arched. Lateral margin of pronotum straight. Posterior lobe of prontum moderately densely punctate with small punctures. Membrane reaching apex of abdomen. Metafemora not more than 1.25 thicker than mesofemora. Mesofemora bearing some tiny tubercles in a row posteroventrally, metafemora with anteroventral row of short stiff setae and a few tubercles posteroventrally. Abdomen finely coriaceous. Tibiae straight.

Measurements (female Holotype): Total body length: 9.50; head: length 1.35, width 1.53, interocular space 0.80; length of antennomeres: I 1.40, II 2.03, III 1.65, IV 1.58; length of labiomeres: I 2.10, II 2.12, III 2.60, IV 1.12; pronotum: length 1.45, width 3.35; scutellum: 1.58 length, width 2.00.

Diagnosis. Orbellis longirostris Zámbó & Kondorosy , sp. nov. differs from all other species except O. madagascariensis ( Slater & O’Donnell, 1999) by the large robust body longer than 8.50 mm and the long labium reaching at least abdominal sternite V. Both species share the straight lateral pronotal margin and the metafemora with strong tubercles in a ventral row. The differences between the two species are as follows: the body of O. madagascariensis is shiny vs. the dull body of the new species; pronotum of O. longirostris has rounded collar vs. V-shaped pronotal collar of O. madagascariensis . The body of O. longirostris is much darker and more variegated than of O. madagascariensis ; the pronotum and scutellum are much darker (on O. madagascariensis only the two submedian spots are fuscous at posterior margin); the ground colour of the hemelytron of O. longirostris is yellow with contrasting dark spots (the veins are also pale) while the hemelytra of O. madagascariensis is almost unicolorous yellowish brown (the veins are white in a few short sections only). The other large species is O. maculosus ( Slater & O’Donnell, 1999) —the only known specimen is 8.54 mm long—which is unique with the large contrasting pale apical part of the corium and the mostly pale posterior half of the pronotum. The coloration of O. longirostris is similar to a typical O. dentifemoralis Zámbó & Kondorosy , sp. nov. or O. polhemi ( Slater & O’Donnell, 1999) but the pattern of the scutellum is different, especially on O. polhemi .

Etymology. The Latin adjective refers to the very long labium.

Remark. Male of this species unknown. Although we decided not to describe new species without male(s) or more specimens, O. longirostris Zámbó & Kondorosy , sp. nov. is the exception because the female holotype is very different from other known females in the genus.

Distribution. Orbellis longirostris Zámbó & Kondorosy , sp. nov. is known only from Central Madagascar, west from the main mountain range ( Fig. 28 ).

Figures

FIGURES 1–11. Dorsal views of Orbellis species: 1. O. typicus Distant, 1913 lectotype; 2. O. polhemi Slater & O’Donnell, 1999; 3. O. maculosus Slater & O’Donnell, 1999 holotype; 4. O. madagascariensis Slater & O’Donnell, 1999; 5. O. calcaratus Zámbó & Kondorosy, sp. nov. holotype; 6. O. dentifemoralis Zámbó & Kondorosy, sp. nov. paratype; 7. O. odonnellae Zámbó & Kondorosy, sp. nov. holotype; 8. O. flavescens Zámbó & Kondorosy, sp. nov. holotype; 9. O. guilberti Zámbó & Kondorosy, sp. nov. holotype; 10. O. nigriapicalis Zámbó & Kondorosy, sp. nov. holotype; 11. O. longirostris Zámbó & Kondorosy, sp. nov. holotype. (Fig. 1 by E. Kondorosy, Fig. 3 E. Guilbert, Figs. 2, 4–11 A. Zámbó).

FIGURES 26–28. Distribution of Orbellis species in Madagascar: 26. previously described Orbellis species (solid symbol: literature data; empty symbol: new data; no new data available for O. maculosus); 27. O. calcaratus and O. dentifemoralis; 28. other new Orbellis species.