Scaptomyza (Grimshawomyia)

O’Grady, Patrick, Bonacum, James, Desalle, R. & Val, Francisca Do, 2003, The placement of Engiscaptomyza, Grimshawomyia, and Titanochaeta, three clades of endemic Hawaiian Drosophilidae (Diptera), Zootaxa 159 (1), pp. 1-16 : 10-12

Scaptomyza (Grimshawomyia)

Diagnosis. The genus Grimshawomyia was described by Hardy (1965) and contains two species, G. palata and G. perkinsi , the latter of which was initially described as a member of the genus Drosophila by Grimshaw (1901). The unique male genitalia, which feature surstyli that are exposed and an epandrium which is developed into a pair of moderately large lateral lobes, extending beyond the apices of the surstyli ( Figs. 5­6 ), suggest that these taxa actually form a clade within the genus Scaptomyza . This clade is also characterized by having the second antennal segment sharply pointed at the apex, extending over the base of the third segment; the vertical and ocellar setae inserted into the somewhat swollen sides of the vertex; a long costal fringe, which extends nearly to vein R 4+5, and the distinctive wing markings.

Included Taxa. Scaptomyza (Grimshawomyia) palata (Hardy) , comb. nov. from Maui and O‘ahu, and S. (Grimshawomyia) perkinsi (Grimshaw) , comb. nov. from Maui, O‘ahu and Hawai‘i ( Nishida, 2002).

KEY TO SPECIES OF THE SUBGENUS Grimshawomyia Hardy

16. Clypeus and lower margin of the face yellow; wing with distinct pattern, apex lightly infuscated; two reclinate orbital setae present on frons; coxae predominantly yellow; foretarsi brown to black; third costal section shorter, roughly 2.5 times longer than fourth; each pleuron with a broad, transverse yellow vitta, lower 1/2 of katepisternum yellow, surstyli longer than wide, each with a row of fine teeth on venter. Hawai‘i ..... ....................................................................................................... perkinsi (Grimshaw)

­ Clypeus pale brown; lower margin of the face predominantly brown, with a thin band of yellow; wing pattern similar to above, but with hyaline area at apex; three distinct reclinate orbital setae present on frons; coxae brown; foretarsi yellow; third costal section longer, about 3.5 times longer than fourth; pleura almost entirely brown, lacking distinct vittae; surstyli plainly visible, evenly rounded on ventral surface, lacking conspicuous teeth. O’ahu, Maui ................................................................... palata (Hardy)

Figures

FIGURES 5­6. 5, Male terminalia of Scaptomyza (Grimshawomyia) perkinsi (Grimshaw); 6, Male terminalia of Scaptomyza (Grimshawomyia) palata (Hardy).