Anemesia, Pocock, 1895

Zonstein, Sergei, 2018, A revision of the spider genus Anemesia (Araneae, Cyrtaucheniidae), European Journal of Taxonomy 485, pp. 1-100 : 8-10

publication ID	https://doi.org/ 10.5852/ejt.2018.485
publication LSID	lsid:zoobank.org:pub:55A0F74D-FA80-4C6A-AD74-B49C9061A449
DOI	https://doi.org/10.5281/zenodo.3848299
persistent identifier	https://treatment.plazi.org/id/03D05632-DF36-9778-CCA8-FCEEFAD70832
treatment provided by	Valdenar
scientific name	Anemesia
status

Treatment

Key to species of Anemesia View in CoL

Males

1. Palpal tibia distinctly swollen and mostly aspinose ( Figs 173–176, 178–179 View Figs 173–187 ) ................................ 2

– Palpal tibia subcylindrical or cylindrical, and generally with numerous spines and thick bristles ( Figs 177, 180–187 View Figs 173–187 ) .......................................................................................................................... 7

2. Embolus shorter and widely tapering ( Figs 188–196 View Figs 188–196 ) ...................................................................... 3 – Embolus longer and narrowly tapering ( Figs 199–205 View Figs 197–205 ) ................................................................... 6

3. Distance AME–AME subequal to diameter of AME ( Figs 48–50 View Figs 48–62 ) .................................................. 4 – Distance AME–AME considerably larger then diameter of AME ( Fig. 51 View Figs 48–62 ) ...................................... .................................................................................................................... A. tubifex ( Pocock, 1889)

4. Carapace and legs light to medium brown; abdomen with distinct dorsal pattern ( Figs 1, 3–4 View Figs 1–9 ) ..... 5 – Carapace and legs dark brown; abdominal pattern indistinct ( Fig. 2 View Figs 1–9 )................................................. ............................................................................ A. koponeni Marusik, Zamani & Mirshamsi, 2014 View in CoL

5. Carapace and legs paler, light brown ( Fig. 3 View Figs 1–9 ); embolus longer ( Figs 193–194 View Figs 188–196 ) .. A. pococki View in CoL sp. nov. – Carapace and legs darker, medium brown ( Fig. 1 View Figs 1–9 ); embolus shorter ( Figs 188–190 View Figs 188–196 ) ........................ ................................................................................................................... A. birulai ( Spassky, 1937) View in CoL

6. Palpal tibia aspinose ( Fig. 179 View Figs 173–187 ). Embolus gradually tapering and uniformly curved ( Figs 203– 205 View Figs 197–205 ) ..................................................................................................................... A. sogdiana View in CoL sp. nov.

– Palpal tibia with numerous spines ( Fig. 178 View Figs 173–187 ). Distal part of embolus more sharply tapering and curved than its proximal part ( Figs 199–202 View Figs 197–205 ) ........................................................ A. oxiana View in CoL sp. nov.

7. Cymbium and/or palpal tibia with spines ( Figs 180–187 View Figs 173–187 ). Embolus long to very long, with slightly curved and gradually tapering distal part ( Figs 206–223 View Figs 206–214 View Figs 215–223 ) ................................................................ 8

– Palpal tibia and cymbium aspinose ( Fig. 177 View Figs 173–187 ). Embolus relatively short and bent medially, with very thin and narrow distal part ( Figs 197–198 View Figs 197–205 ) ....................................................... A. andreevae View in CoL sp. nov.

8. Cymbium with numerous dorsal and few lateral spines ( Figs 180–183 View Figs 173–187 ). Subapical embolic keel very small and gradually elevated ( Figs 206, 208, 210, 213–214 View Figs 206–214 ). PMS small and slender ( Figs 279–280, 282–284 View Figs 278–290 ) ........................................................................................................................................... 9

– Cymbium without lateral spines ( Figs 184–187 View Figs 173–187 ). Subapical embolic keel larger and noticeably dilated ( Figs 147–150 View Figs 143–157 , 215, 218, 220 View Figs 215–223 ). PMS stouter ( Figs 285, 289 View Figs 278–290 ) ............................................ 12

9. Carapace and legs paler, light brownish ( Figs 10–11 View Figs 10–17 ). Palpal tibia shorter: 3.0–3.2 times as long as wide ( Figs 182–183 View Figs 173–187 ). Tarsus IV with numerous (5–20) spines ...................................................... 10

– Carapace and legs darker, chestnut brown ( Figs 8–9 View Figs 1–9 ). Palpal tibia longer: 4.3–4.6 times as long as wide ( Figs 180–181 View Figs 173–187 ). Tarsus IV with few (1–4) spines ...................................................................11

10. Legs longer:tibia I 4.9–5.0 times as long as wide ( Fig. 127 View Figs 125–132 ). Palpal organ pegtop-shaped, with certainly bulging anterior lobe ( Figs 211–212 View Figs 206–214 ). Dorsal abdominal pattern as in Fig. 10 View Figs 10–17 ..... A. pallida View in CoL sp. nov.

– Legs shorter: tibia I 4.2 times as long as wide ( Fig. 128 View Figs 125–132 ). Palpal organ comma-shaped, with weakly convex anterior lobe ( Figs 213–214 View Figs 206–214 ). Dorsal abdominal pattern as in Fig. 11 View Figs 10–17 ..... A. parvula View in CoL sp. nov.

11. Legs longer: tibia I 5.7–5.9 times as long as wide ( Fig. 125 View Figs 125–132 ). Palpal organ with strongly bulging anterior lobe ( Fig. 208 View Figs 206–214 ) ....................................................................................... A. castanea View in CoL sp. nov.

– Legs shorter: tibia I 5.1–5.2 times as long as wide ( Fig. 126 View Figs 125–132 ). Palpal organ with weakly convex anterior lobe ( Fig. 210 View Figs 206–214 ) ...................................................................... A. karatauvi ( Andreeva, 1968) View in CoL

12. Palpal bulb noticeably (1.3–1.4 times) wider than long ( Fig. 215 View Figs 215–223 ). Clypeus wide, about 2 times exceeding diameter of ALE ( Fig. 59 View Figs 48–62 ). Posterior sigilla long, oval, and located closer to sternal margin ( Fig. 101 View Figs 99–107 ) ....................................................................................... A. incana Zonstein, 2001 View in CoL

– Palpal bulb as wider as long, or very moderately (1.1–1.2 times) wider than long ( Figs 218, 220 View Figs 215–223 ). Clypeus considerably narrower, shorter than diameter of ALE, or only slightly exceeding it ( Figs 60– 62 View Figs 48–62 ). Posterior sigilla smaller, rounded or irregular, and located more distantly from sternal margin ( Figs 102–103 View Figs 99–107 ) ................................................................................................................................ 13

13. Legs longer: tibia I 4.8–4.9 times as long as wide ( Fig. 130 View Figs 125–132 ). Embolus more tapering, with base located closer to retrolateral side of palpal organ ( Figs 217–219 View Figs 215–223 ). Apical segment of PLS longer ( Fig. 287 View Figs 278–290 ) ............................................................................................................ A. infumata View in CoL sp. nov.

– Legs shorter: tibia I 4.0–4.3 times as long as wide ( Figs 131–132 View Figs 125–132 ). Embolus less tapering, with base located closer to prolateral side of palpal organ ( Figs 220–223 View Figs 215–223 ). Apical segment of PLS shorter ( Fig. 289 View Figs 278–290 ) ........................................................................................................... A. infuscata View in CoL sp. nov.

Females

Females of A. koponeni Marusik, Zamani & Mirshamsi, 2014 View in CoL and A. parvula View in CoL sp. nov. are unknown.

1. Spermathecae 2–3 branched ( Figs 224–234 View Figs 224–244 )..................................................................................... 2 – Spermathecae entire ( Figs 235–249, 251–265 View Figs 224–244 View Figs 245–265 ) or divided ( Fig. 250 View Figs 245–265 ) .............................................. 5

2. Spermathecae strictly 3-branched, with long stalks and well sclerotised receptacles ( Figs 233–234 View Figs 224–244 ). Habitus as in Fig. 18 View Figs 18–26 ......................................................................................... A. andreevae View in CoL sp. nov.

– Spermathecae mostly 2-branched, with shorter stalks and poorly sclerotised receptacles ( Figs 224– 232 View Figs 224–244 ). Habitus differs ( Figs 15–17 View Figs 10–17 ) .................................................................................................... 3

3. Distance AME–AME considerably larger then diameter of AME ( Fig. 65 View Figs 63–77 ). Spermathecae as in Figs 230–232 View Figs 224–244 . Habitus as in Fig. 17 View Figs 10–17 .......................................................... A. tubifex ( Pocock, 1889)

– Distance AME–AME subequal to diameter of AME ( Figs 63–64 View Figs 63–77 ). Habitus and spermathecae differ ( Figs 15–16 View Figs 10–17 , 224–229 View Figs 224–244 ) ...................................................................................................................... 4

4. Carapace and legs paler, light brown ( Fig. 16 View Figs 10–17 ). Spermathecae as in Figs 228–229 View Figs 224–244 ........................... ............................................................................................................................... A. pococki View in CoL sp. nov.

– Carapace and legs darker, medium brown ( Fig. 15 View Figs 10–17 ). Spermathecae as in Figs 224–227 View Figs 224–244 ................... ................................................................................................................... A. birulai ( Spassky, 1937) View in CoL

5. Spermathecae divided ( Fig. 250 View Figs 245–265 ). Dorsal abdominal pattern reduced to longitudinal median band. Habitus as in Fig. 23 View Figs 18–26 .............................................................................................. A. pallida View in CoL sp. nov.

– Spermathecae entire ( Figs 235–249, 251–265 View Figs 224–244 View Figs 245–265 ). Habitus, including more developed dorsal abdominal pattern, differs ( Figs 19–22, 24–26 View Figs 18–26 ) ................................................................................................. 6

6. Carapace light yellowish brown to medium chestnut brown, chelicerae darker coloured ( Figs 19– 22 View Figs 18–26 ). Spermathecae with relatively long and narrow stalks ( Figs 235–249 View Figs 224–244 View Figs 245–265 ) ..................................... 7

– Carapace and chelicerae uniformly dark brown ( Figs 24–26 View Figs 18–26 ). Spermathecae with relatively short and wide stalks ( Figs 251–265 View Figs 245–265 ) ...................................................................................................... 10

7. Carapace and legs paler, light yellowish brown ( Fig. 20 View Figs 18–26 ). Maxillae with 10–20 cuspules each. Spermathecae with moderately long stalks ( Figs 235–240 View Figs 224–244 ) ............................... A. sogdiana View in CoL sp. nov.

– Carapace and legs darker, foxy to chestnut brown ( Figs 19, 21–22 View Figs 18–26 ). Maxillae with 28–85 cuspules each. Spermathecae with long to very long stalks ( Figs 241–249 View Figs 224–244 View Figs 245–265 ) .................................................. 8

8. Habitus as in Figs 19 View Figs 18–26 , 29 View Figs 27–33 . Dorsal abdominal pattern with dark longitudinal median spot having two large lateral extensions. Spermathecae with more narrowly spaced bases and conical stalks ( Figs 241–244 View Figs 224–244 ). Apical segment of PLS triangular ( Fig. 276 View Figs 266–277 ) .............................. A. oxiana View in CoL sp. nov.

– Habitus as in Figs 21–22 View Figs 18–26 . Dorsal abdominal pattern with smaller lateral extensions. Spermathecae with more broadly spaced bases and cylindrical stalks ( Figs 245–249 View Figs 245–265 ). Apical segment of PLS shortly digitiform (as in Fig. 281 View Figs 278–290 ) ..................................................................................................... 9

9. Clypeus broader, AME and ALE relatively smaller ( Fig. 70 View Figs 63–77 ). Maxillae with 65–70 cuspules each. Spermathecae with narrower spaced, stronger inclined and more tapering receptacles ( Fig. 245 View Figs 245–265 ) ..................................................................................................................... A. castanea View in CoL sp. nov.

– Clypeus narrower, AME and ALE relatively larger ( Fig. 71 View Figs 63–77 ). Maxillae with 40–55 cuspules each. Spermathecae with broader spaced, lesser inclined and less tapering receptacles ( Figs 246– 249 View Figs 245–265 ) .................................................................................................... A. karatauvi ( Andreeva, 1968) View in CoL

10. Clypeus wide, about 2 times exceeding diameter of ALE ( Fig. 73 View Figs 63–77 ). Posterior sigilla long, oval, and located closer to sternal margin ( Fig. 113 View Figs 108–116 ). Spermathecae with widened bases and oblate receptacles ( Figs 251–254 View Figs 245–265 ) ........................................................................................... A. incana Zonstein, 2001 View in CoL

– Clypeus considerably narrower, shorter than diameter of ALE, or only slightly exceeding it ( Figs 74– 77 View Figs 63–77 ). Posterior sigilla smaller, rounded or irregular, and located more distantly from sternal margin ( Figs 114–116 View Figs 108–116 ). Spermathecae differ ( Figs 255–265 View Figs 245–265 )......................................................................11

11. Tarsus IV with 3–4 spines. Spermathecae shorter, cap-like, with poorly defined receptacles ( Figs 255–257 View Figs 245–265 ) ................................................................................................... A. infumata View in CoL sp. nov.

– Tarsus IV with 7–10 spines. Spermathecae longer, mushroom-like, with dilated receptacles ( Figs 258–265 View Figs 245–265 ) ................................................................................................... A. infuscata View in CoL sp. nov.