Anemesia incana Zonstein, 2001

Zonstein, Sergei, 2018, A revision of the spider genus Anemesia (Araneae, Cyrtaucheniidae), European Journal of Taxonomy 485, pp. 1-100 : 39-42

publication ID	https://doi.org/ 10.5852/ejt.2018.485
publication LSID	lsid:zoobank.org:pub:55A0F74D-FA80-4C6A-AD74-B49C9061A449
DOI	https://doi.org/10.5281/zenodo.3848271
persistent identifier	https://treatment.plazi.org/id/03D05632-DF17-9758-CE40-FA8CFAD70C0E
treatment provided by	Valdenar
scientific name	Anemesia incana Zonstein, 2001
status

Treatment

Anemesia incana Zonstein, 2001 View in CoL

Figs 12 View Figs 10–17 , 24 View Figs 18–26 , 32–33 View Figs 27–33 , 35 View Figs 34–41 , 43–44 View Figs 42–47 , 59 View Figs 48–62 , 73 View Figs 63–77 , 84–85 View Figs 78–89 , 101 View Figs 99–107 , 113 View Figs 108–116 , 129 View Figs 125–132 , 138–142 View Figs 133–142 , 147–150, 156–157 View Figs 143–157 , 171–172 View Figs 158–172 , 184 View Figs 173–187 , 215–216 View Figs 215–223 , 251–254 View Figs 245–265 , 285–286 View Figs 278–290 , 302–305, 318–321, 338–344, 363, 373 View Fig

Anemesia incana Zonstein, 2001: 11 View in CoL , figs 1–8 (Ƌ ♀)

Anemesia incana View in CoL – Zonstein 2007: 211, fig. 9 (♀). — Marusik et al. 2014: 3, fig. 9 (Ƌ).

Diagnosis

Anemesia incana differs from the closely related and similar-looking A. infumata sp. nov. and A. infuscata sp. nov. in a considerably wider clypeus and noticeably smaller eyes both in males and females ( Figs 59 View Figs 48–62 , 73 View Figs 63–77 ; cf. Figs 60–62 View Figs 48–62 , 74–77 View Figs 63–77 ). Additionally, males of A. incana are well distinguishable due to their wider palpal organ ( Figs 215–216 View Figs 215–223 ; cf. Figs 217–223 View Figs 215–223 ), and the conspecific females due to their oblate receptacles ( Figs 251–254 View Figs 245–265 ; cf. Figs 255–265 View Figs 245–265 ).

Material examined

Holotype

TAJIKISTAN: Ƌ, foothills of Aruktau Mts , 1 km NE of Ganjina Village, 750 m, 37°58′ N, 68°32′ E, 13 Apr. 1987, S. Zonstein leg. ( AMNH).

GoogleMaps

Paratypes (15 ƋƋ, 15 ♀♀)

TAJIKISTAN: 12 ƋƋ, 9 ♀♀, Aruktau Mts, surroundings of Ganjina, 700–1000 m, 9–13 Apr. 1987, S. Zonstein leg. ( SMNH: 9 ƋƋ, 6 ♀♀; MNHL: 1 Ƌ; ZISP 1 ♀; ZMMU: 2 ƋƋ, 2 ♀♀); 1 Ƌ, 4 ♀♀, same collection data as for preceding but 11 Apr. 1988 ( SMNH); 1 Ƌ, 1 ♀, same collecting data as for preceding but 22 Apr. 1992 ( SMNH).

Additional material (3 ƋƋ, 7 ♀♀)

TAJIKISTAN: 1 ♀, Aruktau Mts, surroundings of Ganjina, 800–950 m, 19 Apr. 1989, S. Zonstein leg. ( SMNH); 1 Ƌ, same collection data as for preceding but 18 Apr. 1992 ( SMNH); 1 Ƌ, 2 ♀♀, same collection data as for preceding but 22 Apr. 2015 ( SMNH); 3 ♀♀, same collection data as for preceding but 27 Apr. 2002, S. Zonstein and A. Feodorov leg. ( SMNH); 1 Ƌ, Panj Karatau Mts, vicinity of Mt Astana, 1600 m, 37°23′ N, 69°15′ E, 23 Apr. 1991, S. Zonstein leg. ( SMNH); 1 ♀, Vahsh Karatau Mts, summit area 4 km NNE of Mt Hojamaston, 1670 m, 38°00.9′ N, 68°58.7′ E, 24 Apr. 2015, S. Zonstein leg. ( SMNH).

Redescription

Male (holotype)

HABITUS. See Fig. 12. View Figs 10–17

MEASUREMENTS. TBL 14.50, CL 6.87, CW 5.63, LL 0.63, LW 1.05, SL 4.25, SW 3.30.

COLOUR. Carapace, chelicerae, sternum, labium, most part of palps and legs intensely dark reddish brown; maxillae, coxae, metatarsi, tarsi and sternal sigilla dark brownish orange; eye tubercle blackened; abdomen dorsally dark grey with brown pattern consisting of wide median longitudinal spot and few paired transverse and slightly inclined short stripes, ventral part of abdomen dark yellowish grey, spinnerets pale yellow.

PROSOMA. Clypeus wide. Eye tubercle as in Fig 59 View Figs 48–62 . Eye diameters and interdistances: AME 0.19(0.27), ALE 0.28, PLE 0.15, PME 0.14, AME–AME 0.24(0.16), ALE–AME 0.22(0.18), ALE–PLE 0.22, PLE– PME 0.08, PME–PME 0.57. Weak cheliceral rastellum consists of about 20 spikes grouped in front of cheliceral fang and on low mound. Each cheliceral furrow with 7 promarginal teeth and 3–4 smaller teeth confined to basal retromargin. Sternum, labium and maxillae as shown in Fig. 101 View Figs 99–107 . Sternal sigilla oval; posterior pair relatively large, long oval, and located distantly from sternal margin. Maxillae with 4–5 cuspules each.

LEGS. Tibia and metatarsus I as shown in Fig. 129 View Figs 125–132 . Scopula distal on metatarsi I and II, very narrowly divided on tarsi I and II, widely divided on tarsus III, mixed, rudimentary, and widely divided on tarsus IV. Trichobothria: 2 rows of 9–10 each on tibiae, 14–17 on metatarsi, 14–17 on tarsi, 8 on cymbium. PTC I–IV with 6–7 teeth on each margin.

SPINATION. Palp: femur d3, p2; patella p2; tibia p3, r2, v3; tarsus d5–6. Leg I: femur d5, pd2; tibia p3, v6–7+m; metatarsus p2, v7. Leg II: femur d5, pd1; patella p3–4; tibia p3, v8; metatarsus p3, v6. Leg III: femur d4–5, pd3, r2; patella p4–5; tibia d1, p3, r3, v10–11; metatarsus d1, p3, r3, v10; tarsus v9–10. Leg IV: femur d5, p2, r3; tibia r3, v9–11; metatarsus p4, r2, v7–8; tarsus v7–10. Patellae I and IV, and tarsi I–II aspinose.

PALP. Tibia, cymbium and palpal organ as shown in Figs 184 View Figs 173–187 , 215–216 View Figs 215–223 . Palpal tibia long, cylindrical and spinose ( Fig. 184 View Figs 173–187 ). Palpal organ with wide asymmetrical bulb and tapering and slightly curved embolus ( Figs 215–216 View Figs 215–223 ).

SPINNERETS. See Fig. 285 View Figs 278–290 . PMS: length 0.63, diameter 0.30. PLS: maximal diameter 0.53; length of basal, medial and apical segments 1.45, 1.07, 0.95; total length 3.47; apical segment shortly digitiform.

Female (paratype)

HABITUS. See Fig. 24. View Figs 18–26

MEASUREMENTS. TBL 17.50, CL 8.15, CW 7.07, LL 0.85, LW 1.63, SL 4.75, SW 3.97.

COLOUR. Similar to that of male, except even darker chelicerae.

PROSOMA. Clypeus and eye tubercle as in Fig 73 View Figs 63–77 . Eye diameters and interdistances: AME 0.23(0.30), ALE 0.28, PLE 0.20, PME 0.18, AME–AME 0.20(0.13), ALE–AME 0.20(0.17), ALE–PLE 0.22, PLE–PME 0.05, PME–PME 0.53. Weak cheliceral rastellum as in male. Each cheliceral furrow with 7–8 promarginal teeth and 5–6 smaller retromarginal teeth. Sternum, labium and maxillae as shown in Fig. 113 View Figs 108–116 . Sternal sigilla as in male. Maxillae with 7–10 cuspules each.

SPINATION. Palp: tibia p5, v18–20; tarsus v1. Leg I: tibia v8–10; metatarsus p1, v10. Leg II: tibia p3, v8–9; metatarsus p3, v8–10. Leg III: patella r2; tibia p2–3, r2, v9–10; metatarsus p3–4, r3, v12–14; tarsus v6–8. Leg IV: tibia r2, v10–12; metatarsus r2, v11–12; tarsus ventrally ca 20. All femora and most part of patellae with long and relatively thin bristles instead of spines; tarsi I and II unarmed.

LEGS. Scopula distal on metatarsi I and II, entire on palpal tarsus, very narrowly divided on tarsus I, narrowly divided on tarsus II, elsewhere absent. Trichobothria: 8–10 in each row on tibiae, 16–20 on metatarsi, 15–18 on tarsi. Palpal claw with 4 promarginal teeth. PTC I–II with 6–7 teeth on each margin. PTC III with 6 teeth on outer margin, 3 teeth on inner margin; PTC IV with 3 teeth and one tooth, respectively.

SPERMATHECAE. Entire with short stout stalks and considerably oblate receptacles ( Fig. 251 View Figs 245–265 ).

SPINNERETS. See Fig. 286 View Figs 278–290 . PMS: length 0.67, diameter 0.50. PLS: maximal diameter 0.93; length of basal, medial and apical segments 1.55, 1.13, 0.95; total length 3.63; apical segment shortly digitiform.

Variation

The length of the carapace varies from 6.30 to 7.92 in males and from 5.96 to 8.43 in females. The colour of the carapace and legs varies from very dark reddish brown to deep blackish-brown. Number of maxillary cuspules varies from 2 to 11.

Habitat

Foothills and low mountain ridges at an altitude of 600–1700 m, covered with ephemerous grasslands and subshrubs (mostly species of Artemisia ), often with tall shrubs or low open forest ( Pistacia vera L., different species of Prunus , Crataegus and Acer ). The typical biotopes preferred by these spiders are shown in Figs 318–321 View Figs 314–321 .

Distribution

South-western Tajikistan ( Fig. 273 View Figs 266–277 ).

Notes

Some illustrations of Anemesia incana used in this study are partially based on the additional (non-type) material. Live spiders are presented in Figs 32–33 View Figs 27–33 , 35 View Figs 34–41 , 43–44 View Figs 42–47 . The dentition of the cheliceral furrow in female and male paratypes is shown in Figs 84 and 85 View Figs 78–89 , respectively. The separated female palps and legs I–IV are shown in Figs 138–142 View Figs 133–142 . The SEM micrographs include those of the palpal organ (147, 149), the tip of the embolus ( Figs 148, 150 View Figs 143–157 ), the tarsal claws ( Figs 156–157 View Figs 143–157 ), the trichobothrial base and the tarsal organ of leg I ( Figs 171 and 172 View Figs 158–172 , respectively), and spigots of PLE ( Fig. 302 View Figs 291–305 ) and PME ( Figs 303–305 View Figs 291–305 ).

AMNH	American Museum of Natural History
SMNH	Department of Paleozoology, Swedish Museum of Natural History
ZISP	Zoological Institute, Russian Academy of Sciences
ZMMU	Zoological Museum, Moscow Lomonosov State University

Copyright notice

No known copyright restrictions apply. See Agosti, D., Egloff, W., 2009. Taxonomic information exchange and copyright: the Plazi approach. BMC Research Notes 2009, 2:53 for further explanation.

Taxonomy

Kingdom	Animalia
Phylum	Arthropoda
Class	Arachnida
Order	Araneae
Family	Cyrtaucheniidae
Genus	Anemesia