Ephutomma Ashmead, 1899

Lelej, Arkady S. & Williams, Kevin A., 2023, Review of the tribe Smicromyrmini Bischoff, 1920 (Hymenoptera: Mutillidae), Zootaxa 5231 (2), pp. 101-140 : 112-114

7. Ephutomma Ashmead, 1899

( Figs 22–25 , 113 )

Ephutomma Ashmead, 1899: 52 , 53, ♁, ♀; Lelej 1985: 183, ♁, ♀; Lelej & Brothers 2008: 22; Brothers & Lelej 2017: 95, ♁, ♀; Pagliano et al. 2020: 172. Junior subjective synonym of Pseudophotopsis André, 1896 according to Brothers 1975: 590. Resurrected as distinct genus and senior objective synonym of Eremomyrme Suárez, 1965 by Lelej & Kabakov 1980: 192.

Eremomyrme Suárez, 1965 .

Type species Mutilla turcestanica Dalla Torre, 1897 , ♁, by automatic designation. Junior objective synonym of Ephutomma Ashmead, 1899 (unnecessary replacement name) according to Lelej & Kabakov 1980: 192.

Type species. Mutilla turcestanica Dalla Torre, 1897 , ♁ (replacement name for Mutilla incerta Radoszkowski, 1877 , ♁, not Spinola, 1841), by original designation.

Diagnosis. MALE. Mandible wide, tridentate, dorsal margin with heighly curved carina, beneath with large subbasal lobe. Clypeus concave, anterior border with medial projection with two teeth or tubercles. Forewing with three radiomedial and two discoidal cells. Stigmatic cell more or less sclerotized. Vosella lamellate, long narrow. FEMALE. Eyes elliptical, approximate, minimal distance between them 1.1–1.3 × longitudinal diameter. Clypeus with medial basal tubercle and curved down apical part. Mesosoma narrow and head somewhat wider than pronotum. T2 with large basal spot of pale setae and posterior band widened medially of same setae; T3 with pale band of pale setae. Pygidial plate elongated, sculptured with smooth apical part.

Diversity and distribution. Seventeen species are recognized in the genus; 10 based on males only, six based on females only, and one known from both sexes. Fifteen species are Palaearctic, E. fletcheri ( Turner, 1911) is Oriental, and E. somalica ( Magretti, 1892) is Afrotropical.

Remarks. Like Dentilla , most species of Ephutomma are apparently nocturnal in behavior, but their body color is generally mostly black. Lelej’s (1985) key includes about half of the currently recognized species.

Figures

FIGURES 17–25. 17–18. Ephucilla naja (Zavattari, 1913), ♁: 17. Lateral habitus, 18. Head, face view; 19–21. E. thalia (Mickel, 1933), ♀, holotype: 19. Dorsal habitus, 20. Lateral habitus, 21. Labels; 22–23. Ephutomma angustata (Skorikov, 1935), ♁: 22. Lateral habitus, 23. Head, face view; 24–25. E. fletcheri (Turner, 1911), ♀: 24. Dorsal habitus, 25. Lateral habitus. (Photographs by K. Williams, 19–21 from Integrated insect types database of Taiwanese species http://twinsecttype.nmns.edu. tw/specimen/?id=NMNS-SDEI-01389).

FIGURES 110–115. Smicromyrmini, ♀, pygidial plate (T6). 110.Andreimyrme rong Williams in Okayasu et al., 2021, holotype; 111. Dentilla curtiventris (André, 1901); 112. Ephucilla thalia (Mickel, 1933); 113. Ephutomma fletcheri (Turner, 1911); 114. Karunaratnea palatupanae Lelej, 2005, paratype; 115. Nemka viduata (Pallas, 1773). (Photographs by K. Williams andA. Lelej (114)).