Stamnodes ceniza, Matson, 2023

Matson, Tanner A., 2023, A review of Mexican Stamnodes (Lepidoptera: Geometridae) with the description of 16 new species, European Journal of Taxonomy 911, pp. 1-79 : 37-42

Stamnodes ceniza sp. nov.

urn:lsid:zoobank.org:act:6174E8AF-DF9C-49EF-8D27-6B312E01E7A3

Figs 17 , 70 , 83 , 94–95

Diagnosis

This taxon is unmistakable and cannot be confused with other known Stamnodes ; the grey predominance of the hindwing underside is especially diagnostic (see Fig. 17b ).

Etymology

The specific epithet, ‘ ceniza ’, Spanish for ash, follows a theme with two other species described in this work: S. favilla sp. nov. (ember or spark) and S. fuego sp. nov. (fire). This name was inspired by the ashen, dark grey apices and grey hindwing underside.

Material examined

Holotype

MEXICO • ♂; Hidalgo , San Agustín Tlaxiaca ; 20.13622° N, 98.81453° W; elev. 2434 msnm; Aug.– Oct. 2021; Elsi B. Pérez-Jarillo leg.; genetic voucher: TAM0325 ; BOLD Process ID: MXGEO096-23; GenBank: OQ507471 ; LEP190410 CNIN.

Paratype

MEXICO • 1 ♀; Querétaro, Chuvejé ; 21°10′41″ N, 99°33′38″ W; 28 Jan. 1998; Balcázar and Ibarra leg.; genetic voucher: TAM0058 ; genitalia: TAM-2020-035 ; CNIN LEP 063243 .

Description

Male

FOREWING LENGTH. 15 mm (n = 1).

HEAD. Antenna filiform and black. Frons dark grey; vertex cream to light orange. Labial palpus short, slightly porrect, subequal to diameter of eye, dark grey and white. Cephalic collar mostly dark grey.

THORAX. Patagium mostly white; tegula mostly light grey. Legs predominantly light grey.

FOREWING. Light orange; costal area dark grey in basal third and with medial trigonate patch; apical area dark grey, this colour extending thinly down outer margin to tornus. Underside more or less concolourous with upperside, perhaps slightly darker, and areas between dark grey patches of costa whiter than above.

HINDWING. Light orange, with large area of dark grey along costa and apical area. Underside predominantly dark grey with subtle off-white patchwork and lacking orange ground colour of above. Darker grey to black ovate patch bordered by off-white scales at tornal area, diameter of patch about half width of hindwing.

ABDOMEN. Grey.

GENITALIA. Abdomen lost from only male specimen.

Female

Outwardly undifferentiated from male.

FOREWING LENGTH. 17 mm (n = 1).

GENITALIA ( Fig. 70 ). Anterior apophysis two-thirds length of posterior apophysis. Ostium large, lamella antevaginalis strongly sclerotized and subcircular. Short and narrow ductus bursae with prominent sclerite flattened on ventral surface and dorso-laterally rolled toward median area. Corpus bursae spherical; bearing two signa, each with inward directed process; one signum situated near posterior base of corpus bursae (near ductus bursae) and second near anterior third, each covered with minute papillae.

Distribution ( Fig. 83 )

Mexico: Stamnodes ceniza sp. nov. is only known from the southern Sierra Madre Oriental localities of the type material.

Biology

The immature stages are unknown. The holotype was collected between August and October 2021, and the paratype from January.

Molecular characterization

Only the holotype of this species has been successfully sequenced. It is represented in BOLD by the BIN: BOLD:AFB2888 (n = 1). The distance to the nearest neighbour, S. disrupta sp. nov. (n = 5), is 4.8%.

Remarks

The abdomen of the male holotype has been lost rendering the description of the male genitalia of this species impossible at this time. Otherwise, the holotype is in decent shape. However, the paratype is not in great shape; the right hindwing, broken and previously pinned underneath the individual, has since been glued back into place, and only a foreleg and midleg remain of its six legs.

Figures

Figs 11–18. Adult Stamnodes Guenée, [1858], dorsal (a) and ventral (b) views. 11. S. fergusoni Matson & Wagner, 2020, holotype, ♂ (USNMENT01771260). 12. S. deceptiva Barnes & McDunnough, 1918 (USNMENT01771261). 13. S. fervefactaria (Grote, 1881) (USNMENT01771262). 14. S. disrupta sp. nov., holotype (BMEC). 15. S. mariachi sp. nov., holotype, ♀ (EMEC174828). 16. S. erupta sp. nov., holotype, ♂ (USNMENT01771245). 17. S. ceniza sp. nov., holotype, ♂ (LEP190410CNIN). 18. S. proana (Druce, 1893), holotype (NHMUK). Scale bar = 1 cm.

Figs 67–70. Female genitalia. 67. S. disrupta sp. nov., paratype (TAM-2023-261, BMEC). 68. S. mariachi sp. nov., paratype (TAM-2023-283, AMNH). 69. S. erupta sp. nov., paratype (TAM- 2023-286, USNM 154212). 70. S. ceniza sp. nov., paratype (TAM-2020-035, CNIN). Scale bar = 2 mm.

Figs 82–89. Distributions of newly described Stamnodes Guenée, [1858]. 82. S. erupta sp. nov. 83. S. ceniza sp. nov.84. S. saltillo sp. nov.85. S. catarina sp. nov. 86. S. churro sp. nov.87. S. ferropulvisa sp. nov. 88. S. calcarea sp. nov. 89. S. tenebrosa sp. nov. Single dots may represent> 1 individual. Scale bars = 400 km.

Fig. 94. COI neighbour-joining tree for Mexican Stamnodes Guenée, [1858] using default Kimura-2P model in the Barcode of Life Project (BOLD). Taxon names followed by BOLD Process ID.

Fig. 95. Biogeographic associations of Mexican Stamnodes Guenée, [1858]. (*) denotes species found in two or more provinces. Underlying biogeographic provincial map taken from Morrone (2020: fig. 4.5) with permission to reproduce this figure granted by author Juan J. Morrone.