Sesieutes lucens Simon, 1897: 500

Dankittipakul & Deeleman-Reinhold, 2013, Delimitation of the spider genus Sesieutes Simon, 1897, with descriptions of five new species from South East Asia (Araneae: Corinnidae), Journal of Natural History 47 (3), pp. 167-195 : 178-180

Sesieutes lucens Simon, 1897

( Figures 1 A,B, 2 A,B, 3 D, 4 A–E, 10 I)

Sesieutes lucens Simon, 1897: 500 , description of male. Deeleman-Reinhold, 2001: 449, figures 734–735.

Type material

Holotype. ♂, Singapore ( MNHN, AR 13829, examined) .

New material

One ♂, Singapore, Bukit Timah Nature Reserve, Jungle Fall Valley , 100 m, rainforest along small stream, 9 June 2001, leg. P.J. Schwendinger ( MHNG, SIM–01 / 01) . 1♂, 2♀, Malaysia, Johor State, Mt Ophir, Puteri Waterfalls , 80–130 m, rainforest along stream, 21–22 May 2004, leg. P.J. Schwendinger ( MHNG, MAL–04 / 03) .

Diagnosis

Males of S. lucens can be recognized by the smooth carapace with few nodules on peripheral margin ( Figure 1A ), ventral hump on palpal femur, broad and membranous apical portion on proximal RTA ( Figures 3 D, 4 A,B), elongated, sharply pointed distal RTA ( Figures 4A, C ), conductor and tegular apophysis originating apically, their apices reaching tip of cymbium ( Figure 4A ). Females can be recognized by the ovoid copulatory orifices ( Figure 4D ), by the shape of transverse proximal ducts and by the tightly coiled insemination ducts with four or five coils ( Figure 4E ).

Redescription

Male (holotype, MNHN) Total length 4.40; prosoma 2.20 long, 1.54 wide; opisthosoma 2.20 long, 1.38 wide. Leg measurements: leg I 6.05 (1.65, 2.20, 1.26, 0.94); II 5.17 (1.38, 1.88, 1.10, 0.82); III 4.38 (1.26, 1.24, 1.10, 0.78); IV 6.71 (1.76, 2.08, 1.88, 0.98). Leg spination. Leg I: Fe p–1, Ti v–2–2 –2–2–2–2–2, Mt v–2–2 –2–2; leg II: Ti pv–1 v–2–2 –2–2–2, Mt v–2–2 –2–2; leg III: Ti p–1 v–2–1 –1 r–1, Mt p–1– 1 v –2 r–1; leg IV: Fe d–1, Ti p–1–1 v–2–2 r–1, Mt p–1, v–1–1 –1, r–1.

Pattern and coloration ( Figure 1A ). Carapace reddish brown; integument smooth, with few nodules on peripheral margin; chelicerae, labium and gnathocoxae orangebrown; sternum brown; legs bicolored, coxae to femora orange, patellae to tarsi yellow. Opisthosoma elongate-ovoid; dorsum pale, without marking; venter pale; dorsal scutum smooth, orange-brown, covering approximately three-quarters length of opisthosoma; epigastric scutum protruding anteriorly, forming grooved collar.

Palp ( Figures 3 D, 4 A–C). Palpal femur with elevated hump on ventral surface. Palpal tibia with short, triangular prolateral excrescence; proximal RTA truncated in ventral view, triangular in prolateral view, with broad and membranous apical surface; distal RTA elongated, sharply pointed, apex triangular, reaching slightly less than half length of tegulum; tegulum elongate-ovoid, strongly excavated apico-prolaterally, with broad sperm duct running obliquely; conductor lanceolated, originating apically, gradually tapering anteriorly; tegular apophysis hyaline, originating near base of conductor.

Female (here designated, MHNG). Total length 4.80; prosoma 2.40 long, 168 wide; opisthosoma 2.80 long, 1.15 wide. Leg measurements: leg I 6.60 (1.80, 2.40, 1.38, 1.02); II 5.64 (1.50, 2.04, 1.20, 0.90); III 4.78 (1.38, 1.36, 1.20, 0.84); IV 7.30 (1.92, 2.28, 2.04, 1.10). Leg spination. Leg I: Fe p–1, Ti v–2–2 –2–2–2–2–2, Mt v–2–2 –2–2; leg II: Ti v– 2–2 –2–2–2–2, Mt v–2–2 –2–2; leg III: Ti p–1–1 v–1–1 –1–1 r–1, Mt p–1 v–1–1 r–1; leg IV: Fe d–1, Ti p–1–1 v–1–1 –1 r–1, Mt p–1, v–1–1, r–1–1.

Pattern and coloration ( Figure 1B ). Carapace brown; integument finely punctated; chelicerae, labium and gnathocoxae dark brown; sternum orange-brown; legs orangebrown, except anterior femora and tibiae dark brown; yellow annulation present on distal half of anterior tibiae. Opisthosoma ovoid; dorsum pale grey, mottled with numerous, minute, pale spots, posteriorly with series of narrowed chevrons; venter pale, without pattern; dorsal scutum semi-circular, heavily sclerotized, orange-brown, occupying quarter of length of opisthosoma.

Genitalia ( Figures 4 D,E, 10 I). Epigastric scutum with pair of ovoid, obliquely situated copulatory orifices; median ducts consisting of anterior, heavily sclerotized coils, and narrower posterior, less sclerotized coils. Internal genitalia with broad and transverse proximal ducts; proximal part of insemination ducts heavily sclerotized, descending posteriorly to weakly sclerotized distal ducts where they form four or five coils then ascending anteriorly, connected to transverse duct and reniform lateral spermathecae.

Variation

The male, which was recently collected in the tropical rainforests of Malaysia, has a slightly shorter distal RTA when compared with the type specimen.

Distribution

Singapore and Malaysia (new record).

Figures

Figure 1. Sesieutes spp., dorsal habitus (A – O). Sesieutes lucens Simon, male (A), female (B). Sesieutes minuatus sp. nov., holotype male (C), paratype female (D). Sesieutes abruptus sp. nov., holotype male (E), paratype female (F). Sesieutes bifidus sp. nov., holotype male (G), paratype female (H). Sesieutes apiculatus sp. nov., holotype male (I), paratype female (J). Sesieutes aberrans sp. nov., holotype male (K). Sesieutes sp. from Vietnam, male (L). Sesieutes sp. from Laos, male (M). Sesieutes borneensis Deeleman-Reinhold, male from Palawan (N). Sesieutes sp. from southern Thailand, female (O). Scale bar 1. 0 mm.

Figure 2. Sesieutes spp. Sesieutes lucens Simon, male, prosoma, ventral view (A); S. lucens Simon, male, ocular region, dorsal view (B); Sesieutes minuatus sp. nov., paratype male, fovea, dorsal view (C); Sesieutes sp., male from Vietnam, fovea, dorsal view (D). Scale bar 0. 2 5 mm.

Figure 3. Sesieutes spp., male palp, retrolateral view (A – C, G, I – L), ventral view (D, E), dorsal view (F, H). Sesieutes bulbosus Deeleman-Reinhold, holotype (A); Sesieutes borneensis Deeleman-Reinhold, holotype (B); Sesieutes apiculatus sp. nov., holotype (C); Sesieutes lucens Simon, holotype (D); Sesieutes abruptus sp. nov., holotype (E, F); Sesieutes scrobiculatus Deeleman-Reinhold, holotype (G, H); Sesieutes aberrans sp. nov., holotype (I); Sesieutes nitens Deeleman-Reinhold, holotype (J); Sesieutes emancipatus Deeleman-Reinhold, holotype (K); Sesieutes minuatus sp. nov., holotype (L). Scale bar 0. 1 mm.

Figure 4. Sesieutes lucens Simon. Holotype, male palp (A – C), ventral view (A), prolateral view (B), retrolateral view (C). Female (D, E), epigyne, ventral view (D); internal genitalia, dorsal view (E). Scale bars 0. 1 mm (A – C); 0. 2 5 mm (D, E).

Figure 10. Sesieutes spp., epigyne, ventral view. Sesieutes minor Deeleman-Reinhold, paratype (A); Sesieutes scrobiculatus Deeleman-Reinhold, paratype (B); Sesieutes nitens Deeleman- Reinhold, paratype (C); Sesieutes bulbosus Deeleman-Reinhold, paratype (D); Sesieutes emancipatus Deeleman-Reinhold, paratype (E); Sesieutes apiculatus sp. nov., paratype (F); Sesieutes bifidus sp. nov., paratype (G); Sesieutes borneensis Deeleman-Reinhold, paratype (H); Sesieutes lucens Simon, female from Malaysia (I).