Dasydorylas, Skevington, 2001

Ramos-Pastrana, Yardany, Marques, Dayse Willkenia A. & Rafael, José Albertino, 2024, Dasydorylas Skevington, 2001 (Pipunculidae, Tomosvaryellini) of Colombia, with description of three new species and a key to Neotropical males, European Journal of Taxonomy 932, pp. 138-157 : 153-154

Key to males of the Neotropical species of Dasydorylas

1. Postpedicel with acuminate apex ( Fig. 29 ); vein M 1 slightly straight or clearly straight ( Fig. 31 ) ........................................................................................................................................................... 2

– Postpedicel with aristiform apex ( Figs 3 , 16 , 42 ); vein M 1 slightly to clearly upward curved ( Figs 5 , 18 , 44 ) ............................................................................................................................................... 7

2. Phallus bifid [fig. 36 in Rafael & Ale-Rocha (2004)] ................................ D. vulcanus Rafael, 2004

– Phallus trifid [ Figs 12 , 25 , 51 ) .......................................................................................................... 3

3. Both surstyli with bases slightly thicker than apices [fig. 25 in Rafael (1991)]; third costal section subequal in length to fourth [fig. 39 in Rafael (1991)] ............................... D. regalis ( Rafael, 1991)

– Both surstyli with bases clearly thicker than apices ( Fig. 34 ); third costal section slightly longer than fourth ( Fig. 31 ) .................................................................................................................................. 4

4. Left surstylus left-directed and right surstylus directed clearly downwards, when seen in frontal view [fig. 35c in Hardy (1943)] ........................................................................... D. cinctus ( Banks, 1915)

– Both surstyli slightly sideways-directed [fig. 100 in Rafael (1995)] or clearly sideways-directed ( Fig. 34 ) ............................................................................................................................................ 5

5. Phallic guide with a tuft of small setae dorsally [fig. 101 in Rafael (1995)]; junction of ventral and frontal margin forming an angle of about 110° [fig. 101 in Rafael (1995)]; phallus trifid, with ejaculatory ducts not distinctly separated [fig. 101 in Rafael (1995)] ..................................................... ................................................................................................................ D. nigripedes ( Hardy, 1954)

– Phallic guide without tuft of setae dorsally ( Fig. 38 ); junction of ventral and frontal margin of phallic guide curved, not forming an angle ( Fig. 38 ); phallus trifid, with ejaculatory ducts distinctly separated only in distal seventh ( Fig. 38 ) ......................................................................................................... 6

6. Both surstyli thickened basally and medially, thin apically, with outer margins sinuous not forming an angle [fig. 54 in Rafael (1995)]; phallic guide with dorsal margin curved upward in distal half [fig. 54 in Rafael (1995)] ............................................................................ D. eremita ( Hardy, 1954)

– Both surstyli with basal half thickened and distal half thin, with outer margin forming an angle ( Fig. 34 ); phallic guide straight, slightly curved upward only in distal third ( Fig. 38 ) ...................... ................................................................................................................... D. nigellus ( Rafael, 1991)

7. Both surstyli more thickened medially than basally and apically, with inner margins sinuous ( Fig. 47 ); phallic guide with upper margin straight and a tuft of small setae basally ( Fig. 51 ) .......... .................................................................................................................. D. santainesensis sp. nov.

– Both surstyli thickened basally and medially, thin apically, with inner margins straight and outer margins curved ( Figs 8 , 21 ); phallic guide with upper margin sinuous and a tuft of small setae centrally ( Fig. 12 ) or with stout and rigid lobe dorsally ( Fig. 25 ) ................................................................... 8

8. Both surstyli with acute apex ( Fig 8 ); phallic guide with upper margins sinuous and a tuft of small setae centrally ( Fig. 12 ); phallus trifid with ejaculatory ducts distinctly separated only in distal quarter ......................................................................................................... D. colombiensis sp. nov.

– Left surstylus with rounded apex, right surstylus with truncated apex ( Fig. 21 ); phallic guide with a stout and rigid lobe dorsally ( Fig. 25 ); phallus trifid with ejaculatory ducts distinctly separated only in distal ninths ........................................................................................................ D. gibber sp. nov.

Figures

Figs 27–39. Dasydorylas nigellus (Rafael, 1991), ♂ (IAvH–M380). 27. Habitus, left lateral view. 28. Habitus, dorsal view. 29. Antenna. 30. Thorax, dorsal view. 31. Wing. 32. Abdomen, dorsal view. 33. Tergites and sternites 6 and 7, ventral view. 34. Terminalia, dorsal view. 35. Left surstylus, lateral view. 36. Right surstylus, lateral view. 37. Hypandrium and gonopods, ventral view. 38. Phallic guide and phallus, left lateral view. 39. Ejaculatory apodeme.

Figs 1–13. Dasydorylas colombiensis sp. nov., ♂, holotype (IAvH–M1063). 1. Habitus, left lateral view. 2. Habitus, dorsal view.3. Antenna.4. Thorax, dorsal view. 5. Wing.6. Abdomen, dorsal view.7. Tergites and sternites 6 and 7, ventral view. 8. Terminalia, dorsal view. 9. Left surstylus, lateral view. 10. Right surstylus, lateral view. 11. Hypandrium and gonopods, ventral view. 12. Phallic guide and phallus, left lateral view. 13. Ejaculatory apodeme.

Figs 14–26. Dasydorylas gibber sp. nov., ♂, holotype (IAvH). 14. Habitus, left lateral view. 15. Habitus, dorsal view. 16. Antenna. 17. Thorax, dorsal view. 18. Wing. 19. Abdomen, dorsal view. 20. Tergites and sternites 6 and 7, ventral view. 21. Terminalia, dorsal view. 22. Left surstylus, lateral view. 23. Right surstylus, lateral view. 24. Hypandrium and gonopods, ventral view. 25. Phallic guide and phallus, left lateral view. 26. Ejaculatory apodeme.

Figs 40–52. Dasydorylas santainesensis sp. nov., ♂, holotype (CEUA–M101599). 40. Habitus, left lateral view. 41. Habitus, dorsal view. 42. Antenna. 43. Thorax, dorsal view. 44. Wing. 45. Abdomen, dorsal view. 46. Tergites and sternites 6 and 7, ventral view. 47. Terminalia, dorsal view. 48. Left surstylus, lateral view. 49. Right surstylus, lateral view. 50. Hypandrium and gonopods, ventral view. 51. Phallic guide and phallus, left lateral view. 52. Ejaculatory apodeme.