Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017

Moctezuma, Victor & Halffter, Gonzalo, 2021, Taxonomic revision of the Phanaeus endymion species group (Coleoptera: Scarabaeidae), with the descriptions of five new species, European Journal of Taxonomy 747, pp. 1-71 : 7

publication ID	https://doi.org/10.5852/ejt.2021.747.1333
publication LSID	lsid:zoobank.org:pub:3F0B6EAF-C616-4865-811A-414A094B590C
DOI	https://doi.org/10.5281/zenodo.4724992
persistent identifier	https://treatment.plazi.org/id/03CA87F2-FFF8-FFC6-FE2E-46A4BEC23B95
treatment provided by	Plazi (2021-04-28 10:25:17, last updated by Jonas 2024-12-06 15:12:01)
scientific name	Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017
status

Treatment

Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 View in CoL

Figs 1A View Fig , 15 View Fig , 18B View Fig , 19B View Fig

Phanaeus bravoensis Moctezuma et al., 2017: 115 View in CoL . View Cited Treatment

Phanaeus bravoensis View in CoL – Kohlmann et al. 2018: 69 View Cited Treatment . — Moctezuma et al. 2019: 253, fig. 5.

Type material

Holotype

MEXCIO • ♂, Moctezuma et al. 2017: 118, figs 6–8 (revised); Guerrero, Chilpancingo de los Bravo ; IEXA.

Type locality

Mexico, Guerrero, Chilpancingo de los Bravo.

Distribution

Mexico, Sierra Madre del sur, central Guerrero ( Fig. 15 View Fig ).

Remarks

This species was recently described by Moctezuma et al. (2017). To the original description we add that the elytral striae are impressed basally as distinct fossae; right lobe of endophallite copulatrix is more developed than left lobe; right lobe of endophallite copulatrix obtusely triangular, arched superiorly; left lobe of endophallite copulatrix lobed, strongly reduced; central ridge more developed than central column ( Fig. 1A View Fig ). For the female, the trituberculate cephalic carina with nearly aligned, similar sized, conical tubercles; frons punctures scarce, almost effaced; pronotal surface distinctly punctate; pronotal process trituberculate with a posterior concavity; all pronotal tubercles rounded, with middle tubercle slightly more developed and projected frontally than lateral tubercles; posterior pronotal midline distinctly impressed.

References

Kohlmann B., Arriaga-Jimenez A. & Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029

Moctezuma V. & Halffter G. 2017. A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae) from Los Chimalapas, Oaxaca, Mexico. The Coleopterists Bulletin 71: 47 - 56. https: // doi. org / 10.1649 / 0010 - 065 X- 71.1.47

Moctezuma V., Deloya C., Sanchez-Huerta J. L. & Halffter G. 2019. A new species of the Phanaeus endymion species group (Coleoptera: Scarabaeidae: Scarabaeinae), with comments on ecology and distribution. Annales de la Societe entomologique de France (N. S.) 55: 249 - 254. https: // doi. org / 10.1080 / 00379271.2019.1577170

Moctezuma V., Sanchez-Huerta J. L. & Halffter G. 2017. Two new species of the Phanaeus endymion species group (Coleoptera, Scarabaeidae, Scarabaeinae). ZooKeys 702: 113 - 135. https: // doi. org / 10.3897 / zookeys. 702.14728

Figures

Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA).

Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta.

Fig. 18. Pronotum of females within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta).C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta).H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002. K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017.

Fig. 19. Elytral integument of species within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez- Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta). H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002.K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017.