Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017

Moctezuma, Victor & Halffter, Gonzalo, 2021, Taxonomic revision of the Phanaeus endymion species group (Coleoptera: Scarabaeidae), with the descriptions of five new species, European Journal of Taxonomy 747, pp. 1-71 : 22-23

publication ID	https://doi.org/10.5852/ejt.2021.747.1333
publication LSID	lsid:zoobank.org:pub:3F0B6EAF-C616-4865-811A-414A094B590C
DOI	https://doi.org/10.5281/zenodo.4726225
persistent identifier	https://treatment.plazi.org/id/03CA87F2-FFEB-FFD7-FE30-44FCBBE13C7B
treatment provided by	Plazi (2021-04-28 10:25:17, last updated by Jonas 2024-12-06 15:12:01)
scientific name	Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017
status

Treatment

Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 View in CoL

Figs 1G View Fig , 15 View Fig , 18H View Fig , 19H View Fig

Phanaeus huichol Moctezuma et al., 2017: 123 View in CoL . View Cited Treatment

Phanaeus huichol View in CoL – Kohlmann et al. 2018: 69 View Cited Treatment , 81, 83, 88. — Moctezuma et al. 2019: 252–253, fig. 5. Phanaeus (Notiophanaeus) huichol View in CoL – Kohlmann et al. 2018: 81 View Cited Treatment .

Type material

Holotype

MEXICO • ♂, Moctezuma et al. 2017: 123 (revised); Jalisco, Mazamitla ; TAMU.

Non-type material revised (18 ♂♂, 20 ♀♀)

MEXICO – Jalisco • 1 ♀; “ Sierra de Talpa (Los Venados CT). 12–15-VIII- 2017. 1570 m. G. Nogueira Col. ”; GHC • 2 ♂♂, 1 ♀; “ Sierra de Talpa. 20°22′35.29″ N, 104°49′8.87″ O, 14/VIII/ 2015. 1380 m. G. Nogueira Col. ”; VMC GoogleMaps • 2 ♂♂, 1 ♀; “ Sierra de Talpa. 20°22′35.29″ N, 104°49′8.87″ O, 22–26/ VIII/ 2013. 1595 m. G. Nogueira Col. ”; VMC GoogleMaps • 2 ♂♂, 1 ♀; “ Tequila, Volcán de Tequila. 20°47′18.03″ N, 103°50′28.99″ O, 25/VIII/ 2012. 1890 m. G. Nogueira Col. ”; VMC GoogleMaps • 2 ♂♂, 1 ♀; “ Tequila, Volcán de Tequila. 20°47′18.03″ N, 103°50′28.99″ O, 24–27/VIII/ 2012. 1665 m. (CT). G. Nogueira Col. ”; VMC GoogleMaps • 1 ♂, 2 ♀♀; “ Telcome, Yolosta. 20°10′40.113″ N, 103°41′25.35″ O, 2/VII/2017. D 1–2. 1845 m. G. Nogueira Col. ”; VMC GoogleMaps • 4 ♂♂, 4 ♀♀; “ El Jacal , 1830 m, 12–14/VIII/94. G. Nogueira col.”; IEXA • 1 ♂, 2 ♀♀; “S. Manatlán , 13/ 16–IX– 95, 1650 m, G. Nogueira col.”; IEXA • 3 ♂♂, 2 ♀♀; “ S. Manatlán , 18/ 20–IX–1995, 1400 m, G. Nogueira col.”; IEXA • 1 ♂, 4 ♀♀; “ S. Manatlán , 18/ 20–IX– 1995, 1650 m, G. Nogueira col.”; IEXA . – Michoacán • 1 ♀; “ 12 km S Uruapan, Cascada Tzararacua. 1450 m. 21-VIII-97 horse dung”; VMC .

Type locality

Mexico, Jalisco, Mazamitla.

Distribution

Sierra Madre Occidental; Jalisco, northern Michoacán, Nayarit and southern Sinaloa ( Fig. 15 View Fig ). The first record of P. huichol from Michoacán is presented herein.

Remarks

The original description of P. huichol indicates that the colour pattern varies from dark metallic green, bright metallic green, to dark green with blue sheen. After revising additional specimens, a bright metallic green morph with red sheen was found by us. To the original description we add the elytral striae impressed basally as distinct fossae; and central ridge and column of endophallite copulatrix similar in size ( Fig. 1G View Fig ). For the female, the head showing a cephalic trituberculate carina with middle tubercle more frontally projected, carinate and less prominent than lateral tubercles; frons with almost effaced punctures; pronotal sculpture completely smooth, with almost effaced punctures; pronotal process trituberculate, with posterior concavity, followed posteriorly by reduced, rounded tubercle; middle tubercle dentiform or rounded, more developed and projected frontally than lateral tubercles; lateral tubercles rounded or carinate; posterior pronotal midline completely effaced.

References

Kohlmann B., Arriaga-Jimenez A. & Ros M. 2018. Dung beetle vicariant speciation in the mountains of Oaxaca, Mexico, with a description of a new species of Phanaeus (Coleoptera, Geotrupidae, Scarabaeidae). ZooKeys 743: 67 - 93. https: // doi. org / 10.3897 / zookeys. 743.23029

Moctezuma V. & Halffter G. 2017. A new species of Phanaeus Macleay (Coleoptera: Scarabaeidae: Scarabaeinae) from Los Chimalapas, Oaxaca, Mexico. The Coleopterists Bulletin 71: 47 - 56. https: // doi. org / 10.1649 / 0010 - 065 X- 71.1.47

Moctezuma V., Deloya C., Sanchez-Huerta J. L. & Halffter G. 2019. A new species of the Phanaeus endymion species group (Coleoptera: Scarabaeidae: Scarabaeinae), with comments on ecology and distribution. Annales de la Societe entomologique de France (N. S.) 55: 249 - 254. https: // doi. org / 10.1080 / 00379271.2019.1577170

Moctezuma V., Sanchez-Huerta J. L. & Halffter G. 2017. Two new species of the Phanaeus endymion species group (Coleoptera, Scarabaeidae, Scarabaeinae). ZooKeys 702: 113 - 135. https: // doi. org / 10.3897 / zookeys. 702.14728

Figures

Fig. 1. Endophallite copulatrix of the Phanaeus endymion species group (except for P. arletteae Arnaud, 2018). A. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (IEXA). B. Phanaeus chiapanecus sp. nov. (TAMU). C. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (IEXA). D. Phanaeus endymion Harold, 1863 (VMC). E. Phanaeus funereus Balthasar, 1939 stat. rev. (VMC). F. Phanaeus halffterorum Edmonds, 1979 (TAMU). G. Phanaeus huichol Moctezuma, Sánchez- Huerta & Halffter, 2017 (IEXA). H. Phanaeus jackenioi sp. nov. (UVGC). I. Phanaeus malyi Arnaud, 2002 (VMC). J. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. (VMC). K. Phanaeus pacificus sp. nov. (UVGC). L. Phanaeus panamensis sp. nov. (TAMU). M. Phanaeus porioni Arnaud, 2001 stat. rev. (VMC). N. Phanaeus pyrois Bates, 1887 (VMC).O. Phanaeus rzedowskii sp. nov. (TAMU).P. Phanaeus zapotecus Edmonds, 2006 (VMC). Q. Phanaeus zoque Moctezuma & Halffter, 2017 (IEXA).

Fig. 15. Distribution of the Phanaeus endymion species group in North and Central America (in part). Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017;P. edmondsi Moctezuma, Deloya & Halffter, 2019; P. halffterorum Edmonds, 1979; and P. huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 by Jose Luis Sánchez-Huerta.

Fig. 18. Pronotum of females within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta).C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta).H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002. K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017.

Fig. 19. Elytral integument of species within the Phanaeus endymion species group. A. Phanaeus arletteae Arnaud, 2018 (modified from Arnaud 2018). B. Phanaeus bravoensis Moctezuma, Sánchez- Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). C. Phanaeus chiapanecus sp. nov. D. Phanaeus edmondsi Moctezuma, Deloya & Halffter, 2019 (by Jose Luis Sánchez-Huerta). E. Phanaeus endymion Harold, 1863. F. Phanaeus funereus Balthasar, 1939 stat. rev. G. Phanaeus halffterorum Edmonds, 1979 (by Jose Luis Sánchez-Huerta). H. Phanaeus huichol Moctezuma, Sánchez-Huerta & Halffter, 2017 (by Jose Luis Sánchez-Huerta). I. Phanaeus jackenioi sp. nov. J. Phanaeus malyi Arnaud, 2002.K. Phanaeus olsoufieffi Balthasar, 1939 stat. rev. L. Phanaeus pacificus sp. nov. M. Phanaeus panamensis sp. nov. N. Phanaeus porioni Arnaud, 2001 stat. rev. O. Phanaeus pyrois Bates, 1887. P. Phanaeus rzedowskii sp. nov. Q. Phanaeus zapotecus Edmonds, 2006. R. Phanaeus zoque Moctezuma & Halffter, 2017.