ARCHITAENIOGLOSSA, Haller, 1890

Dayrat, B. & Tillier, S., 2002, Evolutionary relationships of euthyneuran gastropods (Mollusca): a cladistic re-evaluation of morphological characters, Zoological Journal of the Linnean Society 135 (4), pp. 403-470 : 453

ARCHITAENIOGLOSSA )

Pila Röding, 1798 (Bouvier, 1886b, 1887b, c, 1888; Ranjah, 1942; Andrews, 1964, 1965a, b, 1976a, b; Lutfy & Demian, 1966, 1967; Berthold, 1991; Bieler, 1993). (24) Hypobranchial gland reduced but here considered present. (28) Blood gland surrounding the posterior aorta, and not the anterior one, doubtfully homologized here with euthyneuran blood glands. (31) The nephridial gland is absent in Pila but it is present in other freshwater caenogastropods (Franc, 1968; Andrews, 1976a, b). We code the nephridial gland present because of the occurrence of a vein interpreted as a remnant of the efferent vein of this gland. (32–42) Digestive system similar to that of Marisa cornuarietis (Andrews, 1965b; Lutfy & Demian, 1967; Berthold, 1991). (43–51) A left anastomosis joins the supraoesophageal and left pleural ganglia; a short right anastomosis joins the suboesophageal to the right pleural side by side: {lPl-Sb[rPl]-Abd-Sp[-lPl]- rPl}.

Campanile Bayle, 1884 (Bouvier, 1887a; Houbrick, 1981, 1989). (43–51) Right anastomosis joining the right pleural ganglion and the suboesophageal ganglion (Bouvier, 1887a; Houbrick, 1981, 1989). (60) Glandular part of the inner lamina of the pallial oviduct homologized here with the mucous gland because of its position and its gelatinous secretion (Houbrick, 1981, 1989). (66) Spermatophore probably present (Houbrick, 1989). (67) The spawn is a crescent-shaped mass which includes many capsules surrounded by gelatinous fluid homologous with mucus (Houbrick, 1981). Robertson (1985) coded the chalaze of Campanile as a (?), whereas Haszprunar (1988) considered it present in Campanilimorpha. The capsules, which contain 1– 5 eggs, are joined by a chalaze-like structure which we consider homologous with a true chalaze because it clearly joins capsules and not eggs. The intracapsular fluid is considered to be albumen here, even though it has not been described as such owing of the anatomy of the pallial female glands (Houbrick, 1981).

Cerithium Bruguière, 1789 (Bouvier, 1887c; Johansson, 1953; Marcus & Marcus, 1964; Houbrick, 1971, 1973, 1974a, b, 1988, 1992; Delhaye, 1975; Houston, 1985). (43–51) Nervous system similar to that of Pila except for the very short left visceral cord between the left pleural and the suboesophageal: {lPlSb[-rPl]-Abd-Sp[-lPl]-rPl}. (66) Spermatophores, which are plesiomorphically present in cerithioids (Houbrick, 1971, 1973), are either considered as probably present in the genus Cerithium (Houbrick, 1973) , or exceptionally present in Cerithium muscarum (Houston, 1985) .

Littorina Linné, 1758 (Bouvier, 1887c; Fretter, 1980; Guyomarch-Cousin, 1976; Andrews, 1981, 1988; Reid, 1989, 1996; Fretter & Graham, 1994). (43–51) {Pl-Sb- Abd-Sp-Pl} (Bouvier, 1887c; Fretter & Graham, 1994).

Coriandria Tomlin, 1917 [The genus Cingulopsis Fretter & Patil (1958) has been synonymized to Coriandria by Lebour (1937), Fretter (1953), Fretter & Patil, 1958, and Ponder & Yoo, 1980]. (43–51) {PlSb- AbdSpPl} (Fretter & Patil, 1958). (66) Males aphallic, internal fertilization allowed by a pallial water flow largely created by a ciliated osphradium and a muscular duct from the posterior part of the pallial cavity to the carrefour (Fretter & Patil, 1958).