Oedodactylus

Herman, Lee, 2010, Generic Revision Of The Procirrina (Coleoptera: Staphylinidae: Paederinae: Pinophilini), Bulletin of the American Museum of Natural History 2010 (347), pp. 1-78 : 63-64

procirrus, Oedodactylus , Pseudoprocirrus , and Stylokyrtus .

DESCRIPTION: Head (fig. 6) not pedunculate, elongate, longer than wide, with postocular region tapered to neck; frontoclypeus with uninterrupted, subapical, interantennal, transverse ridge; postocular lateral margin broadly and shallowly rounded and strongly convergent to neck; postocular margin long; basal angle indistinct; basal margin broadly rounded, evident only anterior to nuchal constriction, and without marginal ridge. Neck across nuchal constriction slightly less than two fifths as wide as greatest postocular width of head; nuchal groove shallow; [**nuchal ridge]. 2 Eye length shorter than postocular length of head. Dorsal surface with reticulate punctation anteriorly and umbilicate punctation medially and posteriorly. Ventral surface without postocular groove. Gular sutures separated; sutures most approximate posteriorly. Gula without pubescence. Antennomere 11 elongate, nearly as long as 8–10 combined to slightly longer than 2–10 combined; apex without spiniform pencil of setae. [**Mandibles, denticle]; [**prostheca]. Maxillary palpus with palpomere 4 longer than third, robust, fusiform, and compressed. [**Ligular region]. [**Labium, glossae]. [**Hypopharynx]. Labrum with submedial denticle on anterior margin. [**Epipharynx].

Prothorax (fig. 6) rectanguliform, longer than wide; widest near anterior margin then mildly to weakly constricted just distad of middle then slightly expanded to base. Pronotum with dense, coarse, umbilicate punctation; punctation uniform, surface without midlongitudinal carina, and with short impunctate midlongitudinal strip just proximad of base. Pronotal marginal ridge poorly and irregularly developed, margined dorsally by row of punctures and lateroventrally by impunctate, polished surface. Notosternal suture poorly developed, evident as weak ridge in broad depression; suture and marginal ridge separated. Hypomeron polished and densely punctate. Postprocoxal lobe moderately long and punctate; transverse ridge absent; apex of each lobe only moderately separated from one another medi-

2 This structure and other similarly cited structures were unavailable for study.

ally. Probasisternum without median carina, but with ridge on each side of median tumescence; surface impunctate. Mesospiracular peritreme with anteromedial margin fused to furcasternum. Elytra shorter than pronotum; humeral angles absent; posterior edge without row of setae; subapical region without long, thicker, posteromediad directed seta near apicolateral corner. Scutellum with a few setae. Mesosternum without median carina. [**Mesocoxal acetabulum]. [**Intersternal suture].

Profemur with ridge near middle on anteroventral surface. Protibia with numerous combs [**number of combs] extending for most of length; tibia with shallow depression on ctenidial surface; apical portion neither constricted nor enlarged. Protarsomeres (fig. 6) 1–4 inflated, ventral surface without setose pad; base of tarsomere 1 not surrounded by cupulate protibial apex; tarsomere 4 not expanded beneath tarsomere 5 and apical margin entire, not bilobed; tarsomere 5 unmodified and inserted laterad of middle of asymmetrical tarsomere 4; tarsomere 5 with moderately dense pubescence ventrally. Mesotarsomere and metatarsomere 1 longer than remaining articles combined.

Abdominal segments without imbricate macrosculpturing (cf. fig. 28). Segment III without paratergites; paratergal carina present at base laterad of spiracle; tergum and sternum III fused. Segment VII with tergum and sternum separated. [**Segments IV–VI, ‘‘windows’’]. [**Sternum I]. [**Sternum IV]. Tergum IX fused basally; posterior margin deeply emarginate; lateroapical process short, slender in dorsal view and wider and apically tapered in lateral view, feebly curved ventrally, and extending beyond posterior margin of tergum X; [**base of lateroapical process]. Tergum X with apical margin rounded; base separated from tergum IX.

[**Female genital sclerites].

[**Aedeagus].

DISTRIBUTION AND HABITAT: I examined specimens of P. miricornis from Singapore, the type locality, and G. Rougemont (correspondence, July 2007) collected a specimen from Negri Sembilan, Malaysia, a new record for the species. He collected the Malaysian specimen and another from Singapore by sifting forest floor litter. I examined one specimen of an undescribed species from Malaysia.

DISCUSSION: The genus was originally described as Eucirrus ( Fauvel, 1895: 215) , a preoccupied name replaced with Paraprocirrus ( Bernhauer, 1923) . Fauvel (1895: 216) described the species from one specimen; a second, slightly damaged one is in the Natural History Museum, London, and a third in the collection of Guillaume Rougemont, Londinières, France. Cameron (1928: 439) added P. borneensis , but later moved it to Neoprocirrus ( Cameron, 1936: 42) .

Paraprocirrus miricornis is rarely encountered and known by only three specimens. The fourth antennomere of the species is black while the others are reddish to yellowish brown.

The genus is linked to Neoprocirrus by the elongate terminal antennomere. Fauvel (1895: 216) placed the genus between Procirrus and Oedichirus . The specimens I examined are elongate and slender with long legs and antennae, pale reddish brown, and with coarse, umbilicate punctation on the head and pronotum.

Beyond the descriptions, the localities, and a little bit of collecting data nothing is known about the species.

SPECIES INCLUDED AND MATERIAL EXAMINED

miricornis (Fauvel) – H (IRSN), sp (BMNH, GdRC).............. Malaysia, Singapore (Note: The species appears to have been described from a single specimen. In the original description Fauvel, 1895: 216, wrote: ‘‘Je n’ai vu que l’exemplaire de ma collection,…’’ so that specimen is the holotype ICZN, 1999: Article 73.1.2], not a syntype. In the online list of types in Institut Royal des Sciences Naturelles, Brussels, the type is listed as a lectotype.) UNDETERMINED SPECIMENS: Malaysia.

DISSECTIONS: None.