Phortica (Ashima) foliiseta, Duda (Duda, 1923

Toda, Masanori J., Bänziger, Hans, Sati, Pradeep C., Fartyal, Rajendra S., Suwito, Awit & Katoh, Toru, 2020, Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species, Zootaxa 4789 (1), pp. 1-54 : 12

2. Phortica (Ashima) foliiseta species group, new

Diagnosis. Pregonital, apical knob with fine serrations (ch.45-1; Figs 4C , 5D ) and round barbule (ch.46-0, 47-1; Figs 4C , 5D , 6E , 7G ).

Shared characters. Palpus with hollow-shaped sense organ (ch.9-2). Some anterior, cibarial, medial sensilla short, apically blunt, peg-like (ch.11-1). Pseudotracheae in labellum 6 per side (ch.14-2). Mid and hind femora and hind tibia entirely yellow (ch.16,19-0). Epandrium widely round on ventral margin (ch.31-1; Figs 4B , 5C , 6D , 7E ); setae on and/or near ventral margin of epandrium not denser than those on other portions (ch.32-0; Figs 4B , 5C , 6D , 7E ). Hypandrium symmetric (ch.40-0; Figs 4D , 5E , 6G , 7H ). Pregonite ( Fig. 4C , 5D , 6E , 7G ) partly strongly sclerotized (ch.43-0), rod-like, with apical knob (ch.44-1) and sensilla only on basal portion (ch.53-1) but no serration on ventrodistal portion (ch.50-0). Aedeagal, outer membrane without apical, long, narrow tube (ch.57-0; Figs 5D , 6E , 7G ); aedeagal free sclerite absent (ch.59-0; Figs 5D,E , 6E,G , 7G,H ). Aedeagal sheath and/or postgonites asymmetric (ch.63-1; Figs 4D , 5E , 6F , 7F ).

Included taxa. Phortica (Ashima) foliiseta species subgroup; and P. (A.) tanabei subgroup.

Figures

FIGURE 4. Phortica (Ashima) andreagigoni Toda & Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm.

FIGURE 5. Phortica (Ashima) watabei Sati & Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B.

FIGURE 6. Phortica (Ashima) halimunensis Toda, sp. nov. (A–G; ♂ paratype) and Phortica (Ashima) longipenis Chen & Wen, 2005 (H; ♂ paratype). A, Antenna; B, prementum; C, abdominal sternites; D, periphallic organs (caudolateral view); E, phal- lic organs (lateral view); F, aedeagal sheath and aedeagal median rod (ventral view); G, hypandrium, pregonites and aedeagus (ventral view); H, aedeagus and aedeagal sheath (A-type; ventral view). Scale bars: 0.1 mm in A, B, D–H; 0.5 mm in C.

FIGURE 7. Phortica (Ashima) akutsui Toda, sp. nov. (♂ holotype). A, Head (caudal view); B, frons; C, antenna; D, abdominal sternites; E, periphallic organs (caudolateral view); F, postgonites (anteroventral view); G, H, phallic organs (lateral and ventral views, respectively). Abbreviations: dl t a, dorsolateral, tentorial apodeme; pocl s, postocular setae. Scale bars: 0.1 mm in B, C, E–H; 0.5 mm in A, D.