Phortica (Ashima) foliiseta, Duda (Duda, 1923

Toda, Masanori J., Bänziger, Hans, Sati, Pradeep C., Fartyal, Rajendra S., Suwito, Awit & Katoh, Toru, 2020, Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species, Zootaxa 4789 (1), pp. 1-54 : 15

2.1. Phortica (Ashima) foliiseta species subgroup, new

Diagnosis. Surstylus without peg-like teeth (ch.37-1; Figs 4B , 5C ); pregonite basally with recurved, apically truncate extension (ch.49-3; Figs 4C , 5D ); posterior, rod-like process of phallapodeme longer than aedeagal median rod (ch.62-1; Figs 4C , 5D ).

Shared characters. Prementum without pubescent patches (ch.12-0). Setulae on medial surface of abdominal sternites IV and V not so dense (ch.27,28-0; Fig. 5B ). Setae on and/or near ventral margin of epandrium longer than others (ch.33-1; Figs 4B , 5C ). Aedeagal, outer membrane without spiny ornamentation (ch.55-1; Fig. 5D,E ); aedeagal, median rod less sclerotized (ch.58-2; Fig. 5D ). Aedeagal sheath with lateral lobes bearing strongly sclerotized postgonites (ch.65-1; Figs 4D , 5E ).

Included taxa. Phortica (Ashima) biclavata Chen ; P. (A.) dianmianensis Gong, Zhu & Chen ; P. (A.) foliisetoides Chen & Toda ; P. (A.) libo Zhu, Qin & Chen*; P. (A.) menglian Gong, Zhu & Chen*; P. (A.) nakanoi Chen ; P. (A.) phyllochaeta ; P. (A.) sagittaristula Chen & Wen ; P. (A.) saltiaristula Chen & Wen ; P. (A.) symmetria *; P. (A.) xyleboriphaga *; P. (A.) yalianae Zhu, Qin & Chen*; P. (A.) yiqini Zhu, Qin & Chen*; and P. (A.) foliiseta species complex. * The assignment was judged from the original descriptions by Senior-White (1921), Chen et al. (2005b), Zhu et al. (2017) and Gong et al. (2018).

Figures

FIGURE 4. Phortica (Ashima) andreagigoni Toda & Bänziger, sp. nov. (♂ holotype). A, Antenna; B, periphallic organs (cau- dolateral view); C, aedeagus, phallapodeme, hypandrium and pregonite (lateral view); D, aedeagal sheath, postgonites, hypandrium and pregonites (ventral view). Scale bars: 0.1 mm.

FIGURE 5. Phortica (Ashima) watabei Sati & Fartyal, sp. nov. (♂ paratype). A, Head (frontal view); B, abdominal sternites; C, periphallic organs (caudal view); D, E, phallic organs (lateral and ventral views, respectively; B-type). Scale bars: 0.1 mm in C–E; 0.5 mm in A, B.