Phortica (Ashima) angulata, Prigent & Chen, 2008

Toda, Masanori J., Bänziger, Hans, Sati, Pradeep C., Fartyal, Rajendra S., Suwito, Awit & Katoh, Toru, 2020, Taxonomy and evolution of asymmetric male genitalia in the subgenus Ashima Chen (Diptera: Drosophilidae: Phortica Schiner), with descriptions of seven new species, Zootaxa 4789 (1), pp. 1-54 : 34

4. Phortica (Ashima) angulata species group, new

Diagnosis. Dorsomedial, tentorial apodeme ≥ 2/3 basal, parallel portion of dorsolateral, tentorial apodeme (ch.1-0); abdominal tergites III and IV unicolor (ch.21-3); epandrium with triangular or hook-like projection on anteroventral margin (ch.31-2; Fig. 3B ); pregonite ( Fig. 3C ) not strongly sclerotized (ch.43-2), plate-like (ch.44-0), with basal extension nearly rectangular to main body and apically broad (ch.49-2).

Shared characters. Arista with dorsal but no ventral branches; apical, leaf-like expansion apically with small, awn-like projection. Palpus with hollow-shaped sense organ (ch.9-2). Some anterior, cibarial, medial sensilla short, apically blunt, peg-like (ch.11-1). Prementum without pubescent patches (ch.12-0). Pseudotracheae in labellum 6 per side (ch.14-2). Setulae just before prescutellar acrostichal setae somewhat longer than other acrostichal setulae (ch.15-1). Mid and hind femora and all tibiae entirely yellow (ch.16–19-0). Apical, downward curved setula on fore tarsomere V stout, as long as tarsomere V (ch.20-1). Abdominal sternite III longer than wide (ch.22-0), not wider than IV (ch.23-0); IV not wider than long (ch.24-0); V wider than long (ch.25-1); III–V without dense setulae on medial surface (ch.26–28-0). Setae on and/or near ventral margin of epandrium ( Fig. 3B ) not denser than those on other portions (ch.32-0), longer than others (ch.33-1). Surstylus ( Fig. 3B ) without expansion at caudoventral corner (ch.35-0), with peg-like teeth (ch.37-0). Hypandrium symmetric (ch.40-0); phragma as narrow as lateral gonocoxites (ch.41-1). Pregonite ( Fig. 3C ) not serrated on ventrodistal portion (ch.50-0), with sensilla on submedial and basal portions (ch.52,53-1). Aedeagal, outer membrane bearing neither spiny ornamentation nor apical, long, narrow tube (ch.55-1, ch.57-0); median rod entirely or partly strongly sclerotized (ch.58-0); free sclerite absent within outer membrane (ch.59-0). Posterior, rod-like process of phallapodeme not longer than aedeagal median rod (ch.62-0). Aedeagal sheath with lateral lobes bearing strongly sclerotized postgonites (ch.65-1); sclerotized, posterior postgonites medially separated from each other (ch.66-0).

Included species. This species group comprises African species: Phortica (Ashima) angulata Prigent & Chen ; P. (A.) curvispina Prigent & Chen *; P. (A.) machoruka Prigent & Chen ; and P. (A.) unispina Prigent & Chen *. * The assignment was judged from the original descriptions by Prigent & Chen (2008).

Figures

FIGURE 3. Phortica (Phortica) pappi (Tsacas & Okada, 1983) (A; ♂ from Cape Tribulation, Queensland, Australia), Phortica (Ashima) machoruka Prigent & Chen, 2008 (B, C; ♂ holotype), Phortica (Ashima) efragmentata Toda, sp. nov. (D–G; ♂ ho- lotype) and Phortica (Ashima) afoliolata Chen & Toda, 2005 (H; ♂ paratype from Jianfeng, Ledong County, Hainan, China). A, B, D, Periphallic organs (caudolateral view in A and D; caudal view in B); C, pregonites and hypandrial phragma (ventral view); E, surstylus; F, phallic organs (ventral view); G, H, phallic organs (lateral view). Abbreviations: aed gd, aedeagal guide; aed m r, aedeagal median rod; aed s, aedeagal sheath; cerc, cercus; epand, epandrium; goncx, gonocoxite; hypd ph, hypandrial phragma; phap, phallapodeme; phap pr, posterior rod-like process of phallapodeme; preg, pregonite; postg, postgonite; sur, surstylus. Scale bars: 0.1 mm.